Abstract
A catalog of the types (holotypes, paratypes, lectotypes, paralectotypes) of the species of the family Xylophagidae (Diptera), described by F.D. Pleske (Th. Pleske), N.P. Krivosheina and B.M. Mamaev, and also by V.G. Kovalev, which are stored in the collection of the Zoological Institute of the Russian Academy of Sciences in St. Petersburg, is presented. Photographs of the holotypes and lectotypes and of their labels are given.
All the photographs were taken by N.M. Paramonov using a Canon EOS 800D digital camera with a 65 mm MP-E lens. The images were processed using Helicon Focus 6 software.
The names of the species are listed in alphabetic order and followed by the quoted labels of the type specimens; the current name of the species is given when it differs from the original combination. The labels of the primary types are quoted verbatim, and those of the remaining specimens are given in their complete current or original form in order to make their contents as understandable as possible. Circles of golden paper are pinned under some specimens to designate their belonging to the type series.
The following abbreviations of the names of the depositories are used below:
IEE, A.N. Severtsov Institute of Ecology and Evolution, Moscow, Russia;
ZIN, Zoological Institute, Russian Academy of Sciences, St. Petersburg, Russia;
ZMMSU, Zoological Museum of M.V. Lomonosov Moscow State University, Moscow, Russia.
SPECIES DESCRIBED BY F.D. PLESKE
The identification and some geographical labels were handwritten by F.D. Pleske.
bungei Pleske, 1925 (Pleske, 1925 : 179 (Erinna (Archimyia)) (Fig. 1). The species was described based on 1 male and 5 females without designation of the holotype. The lectotype, male: “17777,” “dol[ina] r[eki] Yany [in Cyrillic, = Yana River valley] [Northern Yakutia], 1885. Dr. A. Bunge,” “no. 20037,” “Er[inna] (Arch[imyia]) bungei Plesk[e] typ.,” “Lectotypus Xylophagus bungei Pl[eske], design. Nartshuk,” designated by E.P. Nartshuk (Krivosheina et al., 1984). The inventory number is INS_DIP_0001182. In the lectotype, largest parts of both wings are missing; the right hind leg and the apex of the abdomen also are missing. The genitalia preparation is stored in a microvial no. 20037.
Paralectotypes. 1 female: “17791” [in “the Green Book” of A.P. Semenov-Tian-Shansky at the Coleoptera Department, the following record corresponds to this number “Dr A. Bunge [?]Choisdah bis Adycha 28.VI– 2.VII”], “[Northern Yakutia] dol[ina] r[eki] Yana [in Cyrillic, = Yana River valley], [?m.] Khitsdakh [Khaiysardakh] and Adycha [? between Khitsdakh and Adycha?] [18]85.VI. (Bunge);” 1 female: “61008,” [Krasnoyarsk Territory] Yamy, N[izhnyaya] Tung[uska] 62°10ʹ 28.VI.1873 (A. Czekanowski);” 1 female: “85846,” “[Yakutia] gran[itsa] lesa na [r.] Golimera [bliz ust’ya r. Olenek] [in Cyrillic, = [Yakutia] forest margin on Golumera River [near Olenek River mouth]] 9.VIII.[1885] (A. Czekanowski);” 1 female: “Sugunur River, verkh[ov’ya] Khara-gola [in Cyrillic, = upper course of Khara-Gol River], Kent [Khentii Mts., Mongolia] 11–12.VII.[1]924 (Kozlov);” 1 female: “Sugu-nur River, verkh[ov’ya] Khara-gola, Kent. [Khentii, Mongolia] 19.VI.[1]924 (Kozlov).” The current name is Xylophagus bungei (Pleske, 1925) (Woodley, 2011).
czerskii Pleske, 1925 (Pleske, 1925 : 182 (Stratioleptis)) (Fig. 2). The species was described based on 3 males and 10 females. The lectotype, male: a golden circle, “Nagornaya, r. Tum[en’-]ula [Tumyn’tszyan], Ross[iisko]-Koreisk[aya] gr[anitsa] [in Cyrillic, = Nagornaya, Tumyn’tszyan River, the North Korea – Russia border] 18.VII.[1]1913 (Czerskii),” “Stratioleptis czerskii Pl. ♂ Typ. Th. Pleske det.,” designated by E.P. Nartshuk (Nartshuk and Kandybina, 1984). The inventory number is INS_DIP_0001183. The lectotype is pinned. The specimen lacks the left middle tarsus; the abdomen is broken off and is glued together with the left hind leg on a separate card pinned under the specimen.
Paralectotypes. 1 female: “okr[estnosti] Slavyanki, Prim[orskaya] o[blast’] [in Cyrillic, = environs of Slavyanka, Primorskaya Province] 23.VI.[1]914 (Emeljanov);” 1 male, 1 female: “r[eka] Tumen-ula, Ross[iisko]-Koreisk[aya] gr[anitsa] [in Cyrillic, = Tyumen-ula River, the North Korea – Russia border] 23.VI.[1]913 (Czerskii);” 3 females: “Nagornaya, Tumen-ula, Ross[iisko]-Koreisk[aya] gr[anitsa] 23.VI.[1]913 (Czerskii);” 1 male, 1 female: “r. Tumen-Ula, Ross[iisko]-Koreisk[aya] gr[anitsa] 27.VI.[1]913 (Czerskii);” 1 female: “Nagornaya, Tumen-ula, Ross[iisko]-Koreisk[aya] gr[anitsa] 18.VII.[1]913 (Czerskii),” “[preparation of genitalia no.] 20043;” 1 female: “Nagornaya, Tumen-ula, Ross[iisko]-Koreisk[aya] gr[anitsa] 26.VII.[1]913 (Czerskii).” The location of two female paralectotypes has not been established.
The current name is Odontosabula czerskii (Pleske, 1925) (Woodley, 2011).
kowarzi Pleske, 1925 (Pleske, 1925 : 178 (Erinna (Archimyia)) (Fig. 3). The species was described based on 1 male. The holotype: a golden circle, “Waldegg [Austria],” “k[ollektsiya] Kovacha [in Cyrillic, = Kowarz’s collection],” “81,” “Er[inna] (Arch[imyia]) kowarzi Plsk. typ.,” “Erinna atra F. Nartshuk det. 1961,” “Holotypus.” The inventory number is INS_DIP_0001184. In the holotype, the right wing, all right legs, left hind leg, 3rd–5th segments of the left fore tarsus, and 2nd–5th segments of the left middle tarsus are missing. The current name is Xylophagus ater Meigen, 1804 (Woodley, 2011).
sachalinensis Pleske, 1925 (Pleske, 1925 : 180 (Erinna (Xylophagus)) (Fig. 4). The species was described based on 1 female. The holotype: a golden circle, “Sakhalin, [without date] D-r [P.I.] Suprunenko,” “Er[inna] (Xyloph[agus] sachalinensis, Plsk. typ.,” “Holotypus.” The inventory number is INS_DIP_0001185. In the holotype, the right wing, right fore and middle legs, left hind leg, the 4th and 5th segments of the left middle tarsus, and 2nd–5th segments of the right hind tarsus are missing. The left wing is partly broken off. The left hind femur and tibia are glued on a separate card. The current name is Xylophagus sachalinensis (Pleske, 1925) (Woodley, 2011).
SPECIES DESCRIBED N.P. KRIVOSHEINA AND B.M. MAMAEV
admirandus Krivosheina et Mamaev, 1972 (Krivosheina and Mamaev, 1972 : 442 (Xylophagus)) (Fig. 5). The species was described based on 10 males and 15 females. The holotype, female: “[Primorskii krai] Suputinskii [Ussuriiskii zapovednik [in Cyrillic, = Primorskii Territory, Suputinskii [Ussuriiskii Nature Reserve]] (216), Amur maackia [Maackia amurensis] 20.V.[19]69 (N. Kr[ivosheina]),” “Holotypus Xylophagus admirandus det. Kriv[osheina] [et] Mam[aev], [19]72.” The inventory number is INS_DIP_0001186. In the holotype, the left and right hind legs and the 2nd–5th segments of the left middle tarsus are missing, otherwise the specimen is well preserved.
Paratypes. 1 female: “Klimoutsy, Amur[skaya] obl[ast’] [in Cyrillic, = Amurskaya Province], 40 km W of Svobodnoe 25.V.[1]957 (Zinov’yev),” “dubrava lespedetsievaya [in Cyrillic, = oak forest with Lespedeza];” 2 males, 1 female: “Suputinskii [zapovednik [in Cyrillic, = Nature Reserve]], (216), akatnik [in Cyrillic, = Amur maackia] 20.5.[19]69 (N. Kr[ivosheina]);” 1 female: “Suputinskii [zapovednik], (86), bird cherry 24.4.[19]69 (N. Kr[ivosheina]);” 1 female: “Suputinskii [zapovednik], (134), walnut tree 5.5.[19]69 (N. Kr[ivosheina]);” 1 female: “Suputinskii [zapovednik], (78), walnut tree 23.4.[19]69 (N. Kr[ivosheina]);” 1 male, 1 female: “k № 48 14.IV.[19]67, Primorsk[ii] kr[ai], Suputinskii z[apovednik] p[od] k[oroi] il’ma [in Cyrillic, = to no. 48 14.IV.[19]67, Primorskii Territory, Suputinskii Nature Reserve, under the elm bark], “k № 48 1967 tam zhe [in Cyrillic, = to no. 48 1967 in the same locality];” 1 male: “Suputinsk[ii] z[apovednik], il’m [in Cyrillic, = Suputinskii Nature Reserve, elm] 21.IX.[19]64 (E. Sogbar). V[ylet] [in Cyrillic, = Emergence] 11.II.[1965];” 1 female: “Suputinskii [Nature Reserve], (147), elm 6.5.[19]69 (N. Kr[ivosheina]);” 1 male: “Suputin[skii] [Nature Reserve], Primorsk[ii] Terr. (27) 6.9.[1]968 ([N.] Krivosheina);” 1 female: “Suputin[skii] z[apovednik], chern[aya] drev[esina] [in Cyrillic, = Suputinskii Nature Reserve, black wood] (36) 13.IV.[19]67;” 1 female: “[zapovednik] Kedrovaya Pad’ [in Cyrillic, = Kedrovaya Pad’ Nature Reserve] (30) 24.VIII.[19]64 V[ylet] [in Cyrillic, = Emergence] 26.II [1965];” 1 female: “Suput[inskii] zap[ovednik], (147-?), elm 15.IX.[19]64 V[ylet] 22.II. Apparently in sample with nos. 151 and 149.” The pupal exuvia are pinned under all the paratypes. The rest 10 paratypes, males and females, are deposited in IEE.
caucasicus Krivosheina et Mamaev, 1982 (Krivosheina and Mamaev, 1982 : 28 (Xylophagus)). The species was described based on 4 males and 4 females. The holotype, male: “Krasnodar Territory, [village] Novo-Prokhladnoe [currently the territory of the Republic of Adygea], from pupa in a beech white stub 1.V.1959 (B. Mamaev)” (IEE).
Paratypes. 1 female: “Krasnodar Territory [at present, in Adygea], [vill.] Guzeripl, larva under fir bark 10.VII.1971 (N.P. Krivosheina),” temporarily is kept in IEE; 2 males, 3 females from “Krasnodar Territory” (probably, also from Adygea which was a part of Krasnodar Territory at that time), and 1 male from Azerbaijan are deposited in IEE.
lukjanovitshi Krivosheina et Mamaev, 1972 (Krivosheina and Mamaev, 1972 : 442 (Xylophagus)) (Fig. 6). The species was described based on 1 male. The holotype: “Az. SSR [Azerbaijan], Lenkor[anskii] r[aio]n, [selo] Divagach [in Cyrillic, = Azerbaijan SSR, Lenkoran District, Divagach vill.] 5.V.[1]933 (Lukjanovitsh),” “Holotypus Xylophagus lukjanovitshi Kriv.[osheina] [et] Mam.[aev], [19]72.” The inventory number is INS_DIP_0001187. In the holotype, the right fore leg is missing; part of this leg apparently has remained on the pin. Otherwise the specimen is well preserved.
signifer Krivosheina et Mamaev, 1972 (Krivosheina and Mamaev, 1972 : 443 (Xylophagus)) (Fig. 7). The species was described based on 4 males and 11 females. The holotype, female: “b/n [without number], Primorsk[ii] kr[ai], Suputinsk[ii] [Ussuriiskii] zap[ovednik] [in Cyrillic, = Primorskii Terr., Suputinskii [Ussuriiskii] Nature Reserve], v yasene [v drevesine] [in Cyrillic = in ash-tree [?wood]] 6.V.[19]69 [(B. Mamaev)],” “Holotypus Xylophagus signifer Kriv[osheina] et Mam[aev], [19]72,” the pupal exuvium is pinned under the specimen. The inventory number is INS_DIP_0001188. In the holotype, the right wing is cracked; the right hind tarsus lacks the 5th segment. Otherwise the specimen is well preserved.
Paratypes. 1 male, 2 females: “Tigrovaya, Suchan[skii] r[aio]n, Uss[uriiskii] kr[ai] [in Cyrillic, = Tigrovaya [?River], Suchanskii District, Ussuriiskii Territory] 10.VI.[1]927 (Stakelberg),” preparation of genitalia no. 20029; 1 male: “b/n [without number] Primorsk[ii] kr[ai], Suputinsk[ii] zap[ovednik] [in Cyrillic, = Primorskii Territory, Suputinskii Nature Reserve] 23.V.[19]69 [collector’s name is not given];” 1 female: “Primorsk[ii] kr[ai], Suputinsk[ii] zap[ovednik] 21.V.[19]69 [collector’s name is not given];” 1 female: “Suputin[skii] z[apovednik], (132), chernaya dreves[ina] [in Cyrillic, = black wood] 1.V.[19]67 [collector’s name is not given];” 1 female: “Suputin[skii] z[apovednik], (133). chernaya dreves[ina] 1.V.[19]67 [collector’s name is not given],” the pupal exuvium is pinned under the specimen; 1 female: “Suputin[skii] zap[ovednik], Primorskii kr[ai] 9.[1]968 (Mamaev),” the pupal exuvium is pinned under the specimen; 2 males, 5 females with similar geographical labels (IEE).
SPECIES DESCRIBED BY V.G. KOVALEV
mongolicus Kovalev, 1982 (Kovalev, 1982 : 1763 (Xylophagus)) (Fig. 8). The species was described based on 3 females. The holotype: “[Mongolia], Ar[a]-Khangaiskii a[imak], Tevshrulekh, no. 1, meadow, 3, 4, 12.VI.1974 [Z.V. Chadaeva],” “Holotypus Xylophagus mongolicus V. Kovalev.” The inventory number is INS_DIP_0001189. In the holotype, the right fore and left middle legs, and also right hind tibia and tarsus are missing. The specimen is glued on a card; the femur and tibia of one of the legs are also glued on the same card.
Paratypes. 1 female with a similar geographical label in the ZIN collection, and 1 female, in the ZMMSU collection.
REFERENCES
Kovalev, V.G., A new species of the dipterous genus Xylophagus (Xylophagidae) from Mongolia, Zool. Zh., 1982, vol. 61, no. 11, p. 1762.
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Nartshuk, E.P. and Kandybina, M.N., Family Coenomyiidae (Diptera), Katalog tipovykh ekzemplyarov kollektsii Zoologicheskogo instituta AN SSSR. Nasekomye dvukrylye (Diptera). Vyp. 2 (Catalogue of the Type Specimens of the Collection of the Zoological Institute, Academy of Sciences of the USSR, Diptera, Issue 2), Kerzhner, I.M., Ed., 1984, Leningrad: Nauka, p. 5.
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Funding
The study of E.P. Nartshuk and G.M. Suleymanova was based on the collection of the Zoological Institute, Russian Academy of Sciences (state project no. 122031100272-3).
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Nartshuk, E.P., Paramonov, N.M. & Suleymanova, G.M. Type Specimens of Awl-Flies (Diptera, Xylophagidae) in the Collection of the Zoological Institute of the Russian Academy of Sciences in St. Petersburg. Entmol. Rev. 103, 698–705 (2023). https://doi.org/10.1134/S0013873823060118
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DOI: https://doi.org/10.1134/S0013873823060118