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Lower Callovian of Kanev Dislocations, Cherkasy Oblast, Ukraine: Ammonites and Stratigraphy

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Abstract

The Jurassic rocks of the area of Kanev glacial dislocations (Central Ukraine) with the Bathonian coastal marine (lagoon) and the Lower Callovian normal marine sediments have been studied, the upper part of which was unknown to the predecessors. The zonal and infrazonal scale of the Lower Callovian of Kanev dislocations are elaborated by ammonites and include four zones, five subzones, and 14 biohorizons: Elatmae Zone (the upper part: P. elatmae and P. vasily biohorizons); Subpatruus Zone (the uppermost part: Ch. saratovensis (G. toricelli) Biohorizon); Koenigi Zone with the Gowerianum (G. metorchum and G. gowerianum biohorizons), Curtilobum (G. indigestum, G. curtilobum, and G. crucifer biohorizons), and Galilaeii (G. galilaeii, s.l. Biohorizon) subzones; Calloviense Zone with the Calloviense (S. khvalynicum and S. kiselevi biohorizons) and Enodatum (C. enodatum planicerclus, C. e. enodatum, and C. e. aeeta biohorizons) subzones. This scale is compared with those of European Russia and Western Europe. It is found that the base of the Proplanulites koenigi Zone, which was distinguished by Karitzky (1887) in the sections of the north Kanev dislocations, biostratigraphically corresponds to the base of the eponymous Zone, which was distinguished by Buckman (1913) in North Yorkshire and used in a standard scale of Europe. The ammonites of the families Cardioceratidae, Kosmoceratidae, Perisphinctidae, and Macrocephalitidae common in the Early Callovian in the East European marine basin are revised and their evolution and biogeography are analyzed. It is shown that, at the beginning of the Callovian, this vast young epicontinental basin was a “pot” of neoendemic evolution of various groups of ammonites, which migrated from Arctic, West European, and West Tethyan biochorems. The western (Dnieper–Donets) marine area of the basin, which also includes the area of Kanev dislocations, was a main way for migration of marine organisms between the West and East European paleobiogeographic provinces. New species are described: Paracadoceras vasily Gulyaev, sp. nov., Sigaloceras fundator Gulyaev, sp. nov., and S. kiselevi Gulyaev, sp. nov.

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Notes

  1. Unfortunately, the first vast collection and significant part of field notes of A.D. Karitzky were lost during a fire in 1883 (Karitzky, 1890, p. 98). He presented a small collection of Kanev fossils to the Geological Committee (now Karpinsky Russian Geological Research Institute, St. Petersburg); it was also examined by S.N. Nikitin and K.A. Tsytovitch.

  2. The scarcely used subfamily Chamoussetiinae was primarily recognized on the basis of onthogenic features of a suture line of the representatives of the genus Chamoussetia R. Douville (Alekseev and Repin, 1989). The evolution of the Middle Jurassic cardioceratids allows us to ascribe the phylogenetically related genera Paracadoceras Crickmay (sensu lato, incl. subgen. Catacadoceras Bodylevsky), Stenocadoceras Imlay, and Cadochamoussetia Mitta to this subfamily. The evolution paths of the predecessors of Chamoussetiinae and jointly evolved Cadoceratinae diverged, probably, at the end of the Bajocian, when inflated spherocone Greencephalites Repin deviated from discocone Arctocephalites Spath (probably, yet from Cranocephalites Spath) (Fig. 16) (Gulyaev, 2009, 2011, 2014; Mitta, 2016; Mitta and Alsen, 2013; Repin et al., 2007; etc.). In the newest taxonomic review (Howarth, 2017), Greencephalites is included in a synonymic of the subgenus Catacadoceras; however, our phylogenetic reconstruction of the Bajocian–Callovian cardioceratids contradicts this conclusion.

  3. Until recently, the representatives of this group were typically considered in the structure of Pseudoperisphinctinae Schindewolf according to (Arkell, 1950; Treatise…, 1957). In the newest taxonomic review (Enay and Howarth, 2019), a poorly characterized typical genus Pseudoperisphinctes Schindewolf is ascribed to synonymic Wagnericeras Buckman in the structure of the subfamily Zigzagiceratinae Buckman.

  4. As seen from Fig. 17,C. surensis is evidently divided into two chronosubspecies/species; however, their nomenclature division is difficult, because a lectotype (Nikitin, 1885, Tab. X(XII), fig. 54) mentioned in (Mitta and Starodubtseva, 1998, p. 7; Mitta, 1999, p. 128) represents damaged inner whorls, which cannot indicate the belonging to the early or late form (subspecies/species).

  5. A morphologically similar form Amm. lamberti crobyloides is known in the Callovian rocks of Southwest Germany (Quenstedt, 1887, Taf. 90, fig. 19; present work, Plate 8, fig. 1). The single specimen, however, is probably lost (Schlegelmilch, 1985) and its stratigraphic position is discussible, which makes problematic the use of the name “Ch. crobyloides,” as was mentioned previously (Gulyaev, 2005а; Gulyaev and Ippolitov, 2013; Gulyaev et al., 2002, 2014).

  6. Hereinafter, the names of the main types of planispiral shells of ammonites (ecomorphs/biomorphs) follow the classical terminology (Treatise…, 1957) and “Westermann triangle” (Westermann, 1996).

  7. In Fig. 17,P. wutachense (Mitta et al., 2015), which was recently defined in the Herveyi Zone of Southwest Germany, morphometrically occupies a position between P. vasily sp. nov. and C. tschernyschewi. At present, however, it is difficult to say whether it belongs to a common East European phyloline or it is a local geographical derivate of P. elatmae (incl. P. suevicum (Callomon, Dietl) that migrated to the Central European basins.

  8. Typically, Gowericeras is considered a subgenus of the genus Kepplerites Neumayr et Uhlig (s.l.); however, phyletically, the successive species of the former represents an individual specific stage in evolution of Kosmoceratids (Figs. 15, 18). This gives grounds to increase the rank of Gowericeras to genus as was primarily considered by Buckman (1921, p. 54; see also Howarth, 2017).

  9. The representatives of Kepplerites s.s. indicated in (Mitta, 2000, 2004b, 2008а; Mitta and Starodubtseva, 2000) from the lower parts of the Elatmae Zone of the Central Volga region, in fact, originate from the Upper Bathonian Infimum Zone (Gulyaev, 2005а, 2013а, 2015d; Kiselev and Rogov, 2007; Mönnig and Dietl, 2017). At present, poorly preserved remains of Kosmoceratids in the complex with Macrocephalites jacquoti (Douville) are reliably known only in the section near the settlement of Prosek (Nizhny Novgorod oblast) in the lower parts of the Elatmae Zone of the Volga region (Gulyaev, 2001а, 2005а, 2015d; Kiselev and Rogov, 2007).

  10. It is likely that G. hildesheimense (Tintant) from the upper parts of the Herveyi Zone of North Germany is a senior subjective synonym or direct descendant of G. russiense (Tintant, 1963; Mönnig and Dietl, 2017). To date, however, it is characterized only by an incomplete core of an adult body chamber, which hampers a full correct comparison.

  11. Nautilus hylas Reinecke is a possible senior synonym of C. enodatum (Reinecke, 1818, Tab. III, Figs. 24, 25), see (Dietl and Mönnig, 2016). The original of this almost unused species, however, is extremely poorly characterized, which makes impossible to identify even its dimorphic affinity.

  12. Kiselev (2001) denotes this chronosubspecies as C. enodatum crispatum Buckman. If we consider that C. enodatum planicerclus and C. enodatum crispatum are subjective synonyms, the name “planicerclus” is valid according to the principle of “first revised” (Mezhdunarodny…, 2004; article 24.2), i.e., J.H. Callomon (1955).

  13. Multiple attempts at division of Macrocephalites into individual genera or subgenera, which were started by Buckman (1909–1930) in the 1920s, were ambiguous and are still waiting for a wide acceptance of the researchers (see Howarth, 2017), although, at any rate, the subgenus division of this diverse group, which lived for a long time (Middle Bathonian–Middle Callovian) on a vast territory (from Indonesia to East Greenland), was required long ago.

  14. A holotype of M. jacquoti (H. Douvillé, 1878, p. 570) [nom. nov. pro Amm. macrocephalus compressus Quenstedt (1847, Taf. 15, Fig. 1)] probably originates from the C. quenstedtiforme (=“C. quenstedti”) Biohorizon in the lower parts of the Callovian of South Germany (Callomon et al., 1989), although the early archaic (with stronger ribbing) representatives of this species appear in the Swabian Jurassic as early as the end of the Bathonian (Dietl, 1994; Dietze and Hostettler, 2015).

  15. The biohorizons in works (Gulyaev et al., 2002; Mitta and Starodubtseva, 1998) are considered according to the previously adopted principles (Gulyaev et al., 2010; Rogov et al., 2012) as provisory and nomenclature not availabile, because they both were given in these publications without indication of the certain stratotypes.

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ACKNOWLEDGMENTS

During the preparation of the work, the authors studied the paleontological collections from the area of Kanev dislocations in many museums of Ukraine and Russia, which are listed in the Introduction. The authors are grateful to staff of these museums for their kind collaboration. Special gratitude goes to V.P. Gritsenko, head of the Department of Geology of the NMNH NAS, Candidate of Geological–Mineralogical Sciences, who provided substantial support for the study of the Kanev collections in the named museum and also enthusiastically participated in the field works. The authors deeply acknowledge the consultations and advice of Professor V.A. Zakharov, Doctor of Geological–Mineralogical Sciences; M.A. Rogov, Doctor of Geological–Mineralogical Sciences (Geological Institute, Russian Academy of Sciences, Moscow); D.N. Kiselev, Doctor of Geological–Mineralogical Sciences (YSPU); Dr. E. Mönnig (Naturkunde-Museum Coburg, Germany); V.V. Mitta, Doctor of Geological–Mineralogical Sciences (Paleontological Institute, Russian Academy of Sciences, Moscow); and Dr. P. Alsen (Geological Survey of Denmark and Greenland, Copenhagen).

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Correspondence to D. B. Gulyaev or A. P. Ippolitov.

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Translated by I. Melekestseva

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Gulyaev, D.B., Ippolitov, A.P. Lower Callovian of Kanev Dislocations, Cherkasy Oblast, Ukraine: Ammonites and Stratigraphy. Stratigr. Geol. Correl. 29, 767–847 (2021). https://doi.org/10.1134/S0869593821070029

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