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ZooKeys 441: 165–181 (2014)
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
doi: 10.3897/zookeys.441.7198
CHECKLIST
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Checklist of the ‘lower Brachycera’ of Finland:
Tabanomorpha, Asilomorpha and
associated families (Diptera)
Jere Kahanpää1, Kaj Winqvist2, heo Zeegers3
1 Finnish Museum of Natural History, Zoology Unit, P.O. Box 17, FI–00014 University of Helsinki, Finland
2 Mikonkatu 3 C 52, FI–20100 Turku, Finland 3 Eikenlaan 24, 3768 EV Soest, the Netherlands
Corresponding author: Jere Kahanpää (jere.kahanpaa@helsinki.i)
Academic editor: J. Salmela | Received 5 February 2014 | Accepted 9 June 2014 | Published 19 September 2014
http://zoobank.org/2F4D6B47-F935-4B88-8B0B-7F832BB12CCA
Citation: Kahanpää J, Winqvist K, Zeegers T (2014) Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha,
Asilomorpha and associated families (Diptera). In: Kahanpää J, Salmela J (Eds) Checklist of the Diptera of Finland.
ZooKeys 441: 165–181. doi: 10.3897/zookeys.441.7198
Abstract
A checklist of the ‘lower Brachycera’ of Finland is presented. his part of the complete checklist of Finnish
Diptera covers the families Acroceridae, Asilidae, Athericidae, Bombyliidae, Mythicomyiidae, Rhagionidae, Scenopinidae, Stratiomyidae, Tabanidae, herevidae, Xylomyidae and Xylophagidae.
Keywords
Species list, Finland, Diptera, biodiversity, faunistics
Introduction
his part of the checklist of the Diptera of Finland covers non-eremoneuran true lies
(Diptera: Brachycera). he brachyceran lies excluded from the clade Eremoneura are
often called the ‘lower Brachycera’ due to their basal position in the true ly tree of life.
It remains unclear whether this assemblage of families is a monophyletic clade. here
are also several models for the relative relationships of the various superfamilies and
families. A simple classiication scheme following Marshall (2012) is adopted for this
Copyright Jere Kahanpää et al. This is an open access article distributed under the terms of the Creative Commons Attribution License (CC BY 4.0),
which permits unrestricted use, distribution, and reproduction in any medium, provided the original author and source are credited.
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Jere Kahanpää et al. / ZooKeys 441: 165–181 (2014)
checklist. Only two infraorders, Tabanomorpha and Asilomorpha, are recognized. he
presentation order of families follows Woodley et al. (2009).
World catalogues have recently been published for Stratiomyidae (Woodley
2001, 2011b), Xylomyidae (Woodley 2011a), Xylophagidae (Woodley 2011c),
Bombyliidae (Evenhuis and Greathead 1999, 2003) and Mythicomyiidae (Evenhuis 2002). he Finnish species were last listed by Kahanpää and Winqvist (2005).
Five species have been added since the last checklist: Haematopota italica Meigen,
1804, Lasiopogon septentrionalis Lehr, 1984, Nemotelus infortunatus Kahanpää,
2010, Xylophagus inermis Krivosheina & Krivosheina, 2000 and Zabrachia tenella
(Jaennicke, 1866) (see Kahanpää 2013, Cannings and Kahanpää 2013, Kahanpää
2010, this paper, and Krivosheina and Rozkošný 1990 respectively). Table 1 summarizes the current family species counts for the world, Europe (based on Fauna
Europaea), and Finland.
Tabanomorpha
he stratiomyoid and xylophagoid lineages are often treated as infraorders (Woodley
et al. 2009).
he soldierlies (Stratiomyidae) are very diverse in the tropics but the species diversity decreases sharply towards the higher latitudes. he wood soldier lies (xylomyids)
is a small ly family associated with dead wood. he Fauna Entomologica Scandinavica
series has a volume on stratiomyoid lies (Rozkošný 1973). A new Nemotelus species
was recently described from Finland (Kahanpää 2010a).
he Finnish rhagionids are relatively well known but a few additional species could
occur in the country. Ptiolina is a problematic genus and the number of recognized
species in Northern Europe has varied from two to seven during the last century.
Athericidae was traditionally placed as a subfamily of Rhagionidae, but it seems more
closely associated with Tabanidae (Marshall 2012). Itämies et al. (1990, 1993) have
studied the distribution of Atherix ibis in Finland.
he Finnish xylophagid fauna is relatively well sampled. Adults of the North European species can be identiied using Nartshuk (1988) or Kahanpää (2009). he larvae
can also be identiied at least at the last larval stage (Stubbs and Drake 2001, Krivosheina and Krivosheina 1966).
he tabanid nomenclature (especially Hybomitra) is quite convoluted and records
in older publications must be taken with a grain of salt. Karvonen (1969) summarized the distribution of tabanids in Finland, but this work is now partially obsolete
due to the diiculties in identifying Hybomitra and Haematopota before Chvála et al.
(1972) was published. For identiication of North European tabanids Chvála et al.
(1972) complemented with pictures in Zeegers and van Haaren (2000) or Krčmar et
al. (2011) is recommended. An illustrated guide to the Finnish species is in preparation
(A. Haarto, unpublished).
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
167
Table 1. Number of species in tabanomorph and asilomorph families plus Acroceridae.
Family
Number of species in
World
Tabanomorpha:
Stratiomyidae
2715 (Woodley 2001, 2011b)
Xylomyidae
138 (Woodley 2011a)
Xylophagidae
134 (Woodley 2011c)
Rhagionidae
694 (Pape et al. 2011)
Athericidae
124 (Pape et al. 2011)
Tabanidae
4405 (Pape et al. 2011)
Asilomorpha:
Asilidae
7513 (Pape et al. 2011)
Bombyliidae
~5000 (Evenhuis and Greathead 1999, 2003)
Mythicomyiidae
~330 (Evenhuis 2002, Pape et al. 2011)
Scenopinidae
416 (Pape et al. 2011)
herevidae
1129 (Pape et al. 2011)
unplaced:
Acroceridae
392 (Pape et al. 2011)
Level of knowledge
Europe
Finland
141
13–14
8
85
10
213
29
1
5
15–16
1
38–39
good
good
average–good
average–good
good
good
524
335
30
17
99
35
18–19
1
3
17
good
good
average
good
average–good
34
5
average
Asilomorpha
he asilids and bombyliids of Finland are rather well known from a faunistic point of view
but little is known about their ecology. Most of the North European species are easy to identify but problems with Villa resulted in a cascade of name changes in the late 20th century.
Falck (2009) and Blöchlinger (2008) are good starting points for identifying Villa adults.
François (1969) has male genitalia igures for some of the more diicult Villa species. he
Mythicomyiidae or micro bee lies were long seen as a subfamily of Bombyliidae.
Identifying hereva species was also fraught with diiculties in the past but by
the end of the 20th century the North European fauna was pretty well understood.
A review of the Finnish therevid fauna with keys has recently been published (Haarto
and Winqvist 2006). he window lies, Scenopinidae, is a smallish asiloid lineage associated with the therevids. It has even been proposed they are a specialized subgroup
of the herevidae (Woodley 2009).
Acroceridae
he small-headed lies (Acroceridae) are a ly family of obscure origin. Ainities with
Nemestrinidae, Tabanoidea, Stratiomyoidea, Bombyliidae and Asilomorpha have been
proposed (see Marshall 2012 for further discussion). Finnish acrocerid records are
mostly of single adults caught by sweep-netting, although Storå (1956) found groups
of 20–40 Acrocera orbiculus swarming on a coastal meadow. he acrocerid species seem
to have declined in abundance during the 20th century. Four of our ive Finnish species are now on the national red list (Kahanpää 2010b).
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Checklist part 1:Tabanomorpha (sensu lato)
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
superfamily Stratiomyoidea Latreille, 1802
STRATIOMYIDAE Latreille, 1802
BERIDINAE Westwood, 1838
BERIS Latreille, 1802
Beris chalybata (Forster, 1771)
Beris clavipes (Linnaeus, 1767)
Beris fuscipes Meigen, 1820
Beris hauseri Stuke, 2004
= strobli auct. nec. Dušek & Rozkošný, 1968
Beris morrisii Dale, 1841
NEMOTELINAE Kertész, 1912
NEMOTELUS Geofroy, 1762
sg. Camptopelta Williston, 1917
Nemotelus nigrinus Fallén, 1817
sg. Nemotelus Geofroy, 1762
Nemotelus infortunatus Kahanpää, 2010
Nemotelus notatus Zetterstedt, 1842
Nemotelus uliginosus (Linnaeus, 1767)
PACHYGASTRINAE Loew, 1856
BERKSHIRIA Johnson, 1914
= Pseudowallacea Kertész, 1921
Berkshiria hungarica (Kertesz, 1921)
= albistylum misid.
= barovskii misid.
NEOPACHYGASTER Austen, 1901
Neopachygaster meromelas (Dufour, 1841)
= orbitalis (Wahlberg, 1854)
ZABRACHIA Coquillett, 1901
Zabrachia minutissima (Zetterstedt, 1838)
Zabrachia tenella (Jaennicke, 1866) see Notes
SARGINAE Walker, 1834
CHLOROMYIA Duncan, 1837
Chloromyia formosa (Scopoli, 1763)
MICROCHRYSA Loew, 1855
Microchrysa cyaneiventris (Zetterstedt, 1842)
Microchrysa lavicornis (Meigen, 1822)
Microchrysa polita (Linnaeus, 1758)
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
SARGUS Fabricius, 1798
Sargus cuprarius (Linnaeus, 1758)
Sargus lavipes Meigen, 1822
= nigripes Zetterstedt, 1842
= splendens auct. nec. Meigen, 1804
Sargus iridatus (Scopoli, 1763)
Sargus ruipes Wahlberg, 1854
STRATIOMYINAE Latreille, 1802
tribe Oxycerini Enderlein, 1914
OXYCERA Meigen, 1803
Oxycera centralis Loew, 1863
= centralis Frey, 1911 preocc.
= freyi Lindner, 1938
Oxycera dives Loew, 1845
Oxycera trilineata (Linnaeus, 1767)
tribe Stratiomyini Latreille, 1802
ODONTOMYIA Meigen, 1803
Odontomyia angulata (Panzer, 1798)
Odontomyia argentata (Fabricius, 1794)
Odontomyia microleon (Linnaeus, 1758)
OPLODONTHA Rondani, 1863
Oplodontha viridula (Fabricius, 1775)
STRATIOMYS Geofroy, 1762
Stratiomys singularior (Harris, 1776)
= furcata Fabricius, 1794
XYLOMYIDAE Verrall, 1901
XYLOMYA Rondani, 1861
Xylomya czekanovskii Pleske, 1925
= interrupta auct. nec. (Pleske, 1926)
= maculata auct. nec. (Meigen, 1804)
superfamily Xylophagoidea Fallén, 1810
XYLOPHAGIDAE Fallén, 1810
XYLOPHAGUS Meigen, 1803
= Erinna Meigen, 1800 suppr.
Xylophagus ater Meigen, 1804 see Notes
= compeditus Wiedemann, 1820
Xylophagus cinctus (De Geer, 1776)
Xylophagus inermis Krivosheina & Krivosheina, 2000 see Notes
= matsumurae misid.
Xylophagus junki (Szilády, 1932)
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Xylophagus kowarzi (Pleske, 1925) see Notes
= ater auct. nec. Meigen, 1804
superfamily Rhagionoidea Latreille, 1802
RHAGIONIDAE Latreille, 1802
RHAGIONINAE Latreille, 1802
RHAGIO Fabricius, 1775
Rhagio annulatus (De Geer, 1776)
Rhagio lineola Fabricius, 1794
Rhagio maculatus (De Geer, 1776)
Rhagio notatus (Meigen, 1820)
Rhagio scolopaceus (Linnaeus, 1758)
Rhagio tringarius (Linnaeus, 1758)
CHRYSOPILINAE Bezzi, 1903
CHRYSOPILUS Macquart, 1826
Chrysopilus auratus (Fabricius, 1805)
?= cristatus (Fabricius, 1775) nom. dubium
Chrysopilus luteolus (Fallén, 1814)
Chrysopilus nubecula (Fallén, 1814)
? Chrysopilus suomianus (Szilády, 1934) see Notes
SPANIINAE Rondani, 1856
OMPHALOPHORA Becker, 1900
Omphalophora oculata Becker, 1900
= lapponica Frey, 1911
PTIOLINA Zetterstedt, 1842
Ptiolina nigra Zetterstedt, 1842
Ptiolina nigrina Wahlgren, 1854 see Notes
Ptiolina nitida Wahlgren, 1854
Ptiolina obscura (Fallén, 1814)
SPANIA Meigen, 1830
Spania nigra Meigen, 1830
SYMPHOROMYIA Frauenfeld, 1867
sg. Paraphoromyia Becker, 1921
Symphoromyia crassicornis (Panzer, 1806)
ATHERICIDAE Nowicki, 1873
ATHERIX Meigen, 1803
Atherix ibis (Fabricius, 1798)
superfamily Tabanoidea Latreille, 1802
TABANIDAE Latreille, 1802
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
CHRYSOPSINAE Lutz, 1905
tribe Chrysopsini Lutz, 1905
CHRYSOPS Meigen, 1803
sg. Chrysops Meigen, 1803
Chrysops caecutiens (Linnaeus, 1758)
Chrysops divaricatus Loew, 1858
Chrysops nigripes Zetterstedt, 1838
= lapponicus Loew, 1858
Chrysops relictus Meigen, 1820
= melanopleurus Wahlberg, 1848
Chrysops ruipes Meigen, 1820
Chrysops sepulcralis (Fabricius, 1794)
Chrysops viduatus (Fabricius, 1794)
= pictus Meigen, 1820
TABANINAE Latreille, 1802
tribe Haematopotini Enderlein, 1922
HAEMATOPOTA Meigen, 1803
Haematopota crassicornis Wahlberg, 1848
Haematopota italica Meigen, 1804
Haematopota pluvialis (Linnaeus, 1758)
= italica misid.
? Haematopota subcylindrica Pandellé, 1883 see Notes
HEPTATOMA Meigen, 1803
Heptatoma pellucens (Fabricius, 1776)
tribe Tabanini Latreille, 1802
ATYLOTUS Osten Sacken, 1876
Atylotus fulvus (Meigen, 1820)
Atylotus plebeius (Fallén, 1817)
Atylotus rusticus (Linnaeus, 1767)
Atylotus sublunaticornis (Zetterstedt, 1842)
HYBOMITRA Enderlein, 1922
Hybomitra arpadi (Szilády, 1923)
Hybomitra astuta (Osten Sacken, 1876) see Notes
= polaris (Frey, 1915)
Hybomitra auripila (Meigen, 1820) see Notes
= aterrima (Meigen, 1820)
Hybomitra bimaculata (Macquart, 1826)
= tropica misid.
?= solstitialis (Meigen, 1820) see Notes
Hybomitra borealis (Fabricius, 1781)
= lapponicus (Wahlberg, 1848)
Hybomitra ciureai (Séguy, 1937)
= schineri Lyneborg, 1959
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Hybomitra distinguenda (Verrall, 1909)
Hybomitra kaurii Chvála & Lyneborg, 1970
= borealis misid.
Hybomitra lundbecki Lyneborg, 1959
= fulvicornis misid.
Hybomitra lurida (Fallén, 1817)
Hybomitra montana (Meigen, 1820)
Hybomitra muehlfeldi (Brauer, 1880)
= laviceps (Zetterstedt, 1842)
Hybomitra nigricornis (Zetterstedt, 1842)
Hybomitra nitidifrons (Szilády, 1914)
= coninis misid.
Hybomitra sexfasciata (Hine, 1923)
= borealis anderi Kauri, 1951
Hybomitra tarandina (Linnaeus, 1758)
Hybomitra tropica (Linnaeus, 1758)
TABANUS Linnaeus, 1758
Tabanus autumnalis Linnaeus, 1761
Tabanus bovinus Linnaeus, 1758
Tabanus bromius Linnaeus, 1758
Tabanus cordiger Meigen, 1820
Tabanus maculicornis Zetterstedt, 1842
Tabanus sudeticus Zeller, 1842
Checklist part 2: Asilomorpha
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
superfamily Asiloidea Latreille, 1802
ASILIDAE Latreille, 1802
ASILINAE Latreille, 1802
ASILUS Linnaeus, 1758
Asilus crabroniformis Linnaeus, 1758
DIDYSMACHUS Lehr, 1996
Didysmachus picipes (Meigen, 1820)
DYSMACHUS Loew, 1860
Dysmachus trigonus (Meigen, 1804)
MACHIMUS Loew, 1849
Machimus setibarbis Loew, 1849
NEOITAMUS Osten Sacken, 1878
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
Neoitamus cothurnatus (Meigen, 1820)
Neoitamus cyanurus (Loew, 1849)
Neoitamus socius (Loew, 1871)
NEOMOCHTHERUS Osten Sacken, 1878
Neomochtherus pallipes (Meigen, 1820)
PAMPONERUS Loew, 1849
Pamponerus germanicus (Linnaeus, 1758)
PHILONICUS Loew, 1849
Philonicus albiceps (Meigen, 1820)
RHADIURGUS Loew, 1849
Rhadiurgus variabilis (Zetterstedt, 1838)
TOLMERUS Loew, 1849
Tolmerus atricapillus (Fallén, 1814)
Tolmerus pyragra (Zeller, 1840)
LAPHRINAE Macquart, 1838
tribe Andrenosomatini Hull, 1962
ANDRENOSOMA Rondani, 1856
Andrenosoma albibarbe (Meigen, 1820)
tribe Laphrini Macquart, 1838
CHOERADES Walker, 1851
Choerades fuliginosus (Panzer, 1798)
Choerades gilvus (Linnaeus, 1758)
Choerades igneus (Meigen, 1820)
Choerades lapponicus (Zetterstedt, 1842)
Choerades marginatus (Linnaeus, 1758)
LAPHRIA Meigen, 1803
Laphria lava (Linnaeus, 1761)
Laphria gibbosa (Linnaeus, 1758)
LEPTOGASTRINAE Schiner, 1862
LEPTOGASTER Meigen, 1803
Leptogaster cylindrica (De Geer, 1776)
Leptogaster guttiventris Zetterstedt, 1842
STENOPOGONINAE Hull, 1962
tribe Dioctriini Hendel, 1936
DIOCTRIA Meigen, 1803
Dioctria atricapilla Meigen, 1804
Dioctria cothurnata Meigen, 1820
Dioctria hyalipennis (Fabricius, 1794)
Dioctria oelandica (Linnaeus, 1758)
Dioctria ruipes (De Geer, 1776)
tribe Stegopogonini Hull, 1962
CYRTOPOGON Loew, 1847
Cyrtopogon lavimanus (Meigen, 1820)
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Cyrtopogon lapponicus (Zetterstedt, 1838)
Cyrtopogon lateralis (Fallén, 1814)
Cyrtopogon luteicornis (Zetterstedt, 1842)
= luteicornis var. pollinosus Frey, 1911
Cyrtopogon pulchripes Loew, 1871
tribe Stichopogonini Hardy, 1930
LASIOPOGON Loew, 1847
Lasiopogon cinctus (Fabricius, 1781)
Lasiopogon septentrionalis Lehr, 1984
BOMBYLIIDAE Latreille, 1802
PHTHIRIINAE Becker, 1913
tribe Phthiriini Becker, 1913
PHTHIRIA Meigen, 1803
Phthiria pulicaria (Mikan, 1796)
BOMBYLIINAE Latreille, 1802
tribe Bombyliini Latreille, 1802
BOMBYLIUS Linnaeus, 1758
sg. Bombylius Linnaeus, 1758
Bombylius discolor Mikan, 1796
Bombylius major Linnaeus, 1758
Bombylius minor Linnaeus, 1758
= allibarbis Zetterstedt, 1842
= albibarbis emend.
SYSTOECHUS Loew, 1855
Systoechus ctenopterus (Mikan, 1796)
= sulphureus (Mikan, 1796)
Systoechus gradatus (Wiedemann, 1820)
ANTHRACINAE Latreille, 1804
tribe Anthracini Latreille, 1804
ANTHRAX Scopoli, 1763
Anthrax anthrax (Schrank, 1781)
Anthrax trifasciatus Meigen, 1804
= leucogaster Wiedemann, 1820
Anthrax varius Fabricius, 1794
tribe Exoprosopini Becker, 1913
EXOPROSOPA Macquart, 1840
Exoprosopa capucina (Fabricius, 1781)
MICOMITRA Bowden, 1964
Micomitra stupida (Rossi, 1790)
tribe Villini Hull, 1973
HEMIPENTHES Loew, 1869
Hemipenthes maura (Linnaeus, 1758)
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
Hemipenthes morio (Linnaeus, 1758)
THYRIDANTHRAX Osten Sacken, 1886
hyridanthrax fenestratus (Fallén, 1814)
VILLA Lioy, 1864
Villa cingulata (Meigen, 1804)
? Villa halteralis (Kowarz, 1883) see Notes
Villa hottentotta (Linnaeus, 1758)
Villa modesta (Meigen, 1820)
Villa occulta (Wiedemann, 1820)
MYTHICOMYIIDAE Melander, 1902
GLABELLULINAE Cockerell, 1914
GLABELLULA Bezzi, 1902
Glabellula arctica (Zetterstedt, 1838)
SCENOPINIDAE Burmeister, 1835
SCENOPINUS Latreille, 1802
Scenopinus fenestralis (Linnaeus, 1758)
Scenopinus niger (De Geer, 1776)
Scenopinus sp. A see Notes
= vitripennis misid.
THEREVIDAE Newman, 1834
THEREVINAE Newman, 1834
ACROSATHE Irwin & Lyneborg, 1981
Acrosathe annulata (Fabricius, 1805)
DIALINEURA Rondani, 1856
Dialineura anilis (Linnaeus, 1761)
DICHOGLENA Irwin & Lyneborg, 1981
Dichoglena nigripennis (Ruthe, 1831)
PANDIVIRILIA Irwin & Lyneborg, 1981
Pandivirilia eximia (Meigen, 1820)
PSILOCEPHALA Zetterstedt, 1838
Psilocephala imberbis (Fallén, 1814)
SPIRIVERPA Irwin & Lyneborg, 1981
Spiriverpa lunulata (Zetterstedt, 1838)
= clausa (Frey, 1911)
THEREVA Latreille, 1796
hereva cinifera Meigen, 1830
= subfasciata Schummel, 1830
hereva fuscinervis Zetterstedt, 1838
hereva handlirschi Kröber, 1912
= praestans Collin, 1948
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hereva inornata Verrall, 1909
hereva lanata Zetterstedt, 1838
hereva microcephala Loew, 1847
hereva nobilitata (Fabricius, 1775)
hereva plebeja (Linnaeus, 1758)
hereva strigata (Fabricius, 1794)
hereva unica (Harris, 1780)
= bipunctata Meigen, 1820
hereva valida Loew, 1847
= circumscripta auct. nec. Loew, 1847
Checklist part 3: families of uncertain position (incertae sedis)
suborder Brachycera Macquart, 1834
clade Orthorrapha Brauer, 1863
? superfamily Nemestrinoidea Griith & Pidgeon, 1832
ACROCERIDAE Leach, 1815
ACROCERA Meigen, 1803
= Paracrocera Mik, 1886
sg. Acrocera Meigen, 1803
Acrocera orbiculus (Fabricius, 1787)
= globulus (Panzer, 1804)
= borealis Zetterstedt, 1838
OGCODES Latreille, 1796
sg. Ogcodes Latreille, 1796
Ogcodes borealis Cole, 1919 see Notes
Ogcodes gibbosus (Linnaeus, 1758)
Ogcodes nigripes (Zetterstedt, 1838) see Notes
Ogcodes pallipes Latreille in Olivier, 1812
Excluded species
Anastoechus nitidulus (Fabricius, 1794) labeling mistake
Beris geniculata Curtis, 1830 misidentiied
Cliorismia ardea (Fabricius, 1794) not found within present borders
Cliorismia rustica (Panzer, 1804) not found within present borders
Choerades imbriata (Meigen, 1820) mistake
Choerades ursulus (Loew, 1851) misidentiied see Notes
Chrysopilus splendidus (Meigen, 1820) mistake
Cyrtopogon maculipennis (Macquart, 1834) labeling mistake
Checklist of the ‘lower Brachycera’ of Finland: Tabanomorpha, Asilomorpha...
177
Epitriptus arthriticus (Zeller, 1840) mistake
Machimus gonatistes (Zeller, 1840) not found within present borders
Odontomyia hydroleon (Linnaeus, 1758) not found within present borders
Pandivirilia nigroanalis (Kröber, 1928) misidentiied
Phthiria canescens Loew, 1846 not found within present borders
Tabanus miki Brauer, 1880 misidentiied
Tolmerus cingulatus (Fabricius, 1781) mistake
Villa fasciata (Meigen, 1804) not found within present borders
= circumdata (Meigen, 1820)
= venusta (Meigen, 1820)
Villa longicornis Lyneborg, 1965 not found within present borders
Villa panisca (Rossi, 1790) not found within present borders
= circumdata auct. nec. (Meigen, 1820)
Xylophagus matsumurae Miyatake, 1965 misidentiied
Notes
Choerades ursulus (Loew, 1851) is a poorly known taxon. It was synonymized with
C. fuliginosus by Lehr (1991) but later considered valid by Bosák and Hradský
(2001). Kahanpää and Winqvist (2005) accepted it as a Finnish species but upon
re-examination we consider it most likely that the single Finnish specimen previously identiied as C. ursulus is a dark male of C. fuliginosus.
Chrysopilus suomianus (Szilády, 1934). he type locality of this species is Enontekiö,
Finland (Szilády 1934). Unfortunately the type material seems lost and the name
is probably best treated as a nomen dubium. Based on Szilády’s original description
it may be a dark form of C. nubecula.
Hybomitra astuta (Osten Sacken, 1876). Kahanpää and Winqvist (2005) could not
locate any material in Finnish collections. Several new records of this species have
since been made and its presence in Finland is now conirmed.
Hybomitra auripila (Meigen, 1820). he synonymy of Hybomitra auripila (Meigen,
1820) with H. aterrima (Meigen, 1820) was established by Schacht (1994) and is
accepted here. Schiner (1862) already mentioned H. aterrima as synonym to H.
auripila. Since we consider him to be the irst revisor, the name H. auripila is valid
under the current Code.
Hybomitra solstitialis (Meigen, 1820) has long been known to be a problematic taxon. It is separated from H. bimaculata (Macquart, 1826) based on color characters
alone. he examined Finnish material includes a range of intermediates between
typical H. bimaculata and H. solstitialis forms. It seems likely that the two names are
synonymous, but types should be consulted before synonymy is formally published.
Haematopota subcylindrica Pandellé, 1883. First recorded from Finland by Vuorimies (1984). Unfortunately the specimens listed in his paper could not be found
and their identiication remains somewhat doubtful.
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Ogcodes borealis Cole, 1919. A single Finnish specimen collected in the mid-19th
century is the sole Palearctic record of this species. Originally identiied and published by Hackman (1970), the record was later conirmed by Kahanpää and Winqvist (2005). O. borealis Cole sensu Schlinger (1960) may be a species complex.
Ogcodes nigripes (Zetterstedt, 1838) is probably a senior synonym of O. zonatus
Erichson, 1840.
Ptiolina nigrina Wahlgren, 1854 may be a synonym of P. nigra Zetterstedt, 1842.
Scenopinus sp. A is an apparently undescribed species near S. fenestralis with black
femora. It occurs widely in Finland in association with bird nests.
Villa halteralis (Kowarz, 1883). See Kahanpää and Winqvist (2005) for a discussion
of the single supposed Finnish record of this species.
Xylophagus ater Meigen, 1804. his name has widely been used for two species. Old
Finnish checklists (Frey et al. 1941, Hackman 1980) followed the model also used
in the world checklist Woodley (2011c) and used this name for the species also
known as Xylophagus kowarzi (Pleske, 1925). On the British Isles the name X. ater
is used as a senior name for X. compeditus Wiedemann in Meigen, 1820. According to Alexander and Clements (1991) and Chandler (1998a, b) the British usage
is correct and it is followed here. hus, X. ater is the common species with females
easily identiied by the three stripes of dusting on the mesonotum.
Xylophagus inermis Krivosheina & Krivosheina, 2000 was described as a subspecies
of X. matsumurae Miyatake, 1965 = maculatus Matsumura, 1916 (preoccupied by
X. maculatus Meigen, 1804) (Krivosheina and Krivosheina 2000). It was raised to
a full species status in the recent world catalogue (Woodley 2011c). All collected
Finnish specimens formerly identiied as X. matsumurae were examined and they
belong to X. inermis.
Zabrachia tenella (Jaennicke, 1866). First recorded from Finland by Krivosheina
and Rozkošný (1990). We have examined the Finnish Zabrachia material and
conirmed the presence of both Z. tenella and Z. minutissima in the country.
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