World Catalog and Conspectus on the Family Dryomyzidae (Diptera ...

MYIA, WORLD 12:207–233, CATALOG fig. AND 1. CONSPECTUS ON THE FAMILY DRYOMYZIDAE 2011 207 

<strong>World</strong> <strong>Catalog</strong> <strong>and</strong> <strong>Conspectus</strong> on the Family 

Dryomyzidae (Diptera: Schizophora) 

Wayne N. Mathis & Masahiro Sueyoshi 

Introduction 

The family Dryomyzidae currently includes 25 species that are classified 

into eight genera. Two of these genera, Palaeotimia Meunier <strong>and</strong> Prodryomyza 

Hennig, are monotypic <strong>and</strong> are represented by fossils only, <strong>and</strong> a third 

genus, Dryomyza Fallén, contains two additional fossil species. Several 

Fig. 1: Dryomyza formosa (Wiedemann). 

207

208 MATHIS <strong>and</strong> SUEYOSHI 

genera that were once placed in the Dryomyzidae have been transferred 

to other families, mostly within the superfamily Sciomyzoidea (Coelopidae, 

Helcomyzidae, Helosciomyzidae, Heterocheilidae, Platystomatidae), 

<strong>and</strong> at one time, the family was treated as a taxon within the Sciomyzidae. 

Adults are relatively large for acalyptrates, with body lengths of up to 18 

mm, <strong>and</strong> most specimens are generally yellowish brown or dark gray. The 

distribution of the genera <strong>and</strong> species is restricted to the northern hemisphere, 

<strong>and</strong> the family is noticeably lacking from the tropics or subtropics. 

Only four species are known from the Oriental Region. 

Zoological catalogs, checklists, <strong>and</strong> equivalent databases are indispensable 

tools for anyone needing a reference to a currently accepted name 

<strong>and</strong> frequently to other information relating to that taxon, such as bibliographic 

<strong>and</strong> distributional data. This is possible because most information 

is filed under a species’ scientific name, which is the key to retrieval of information 

from the literature. The system is dynamic, however, <strong>and</strong> subject 

to interpretation. The taxonomic literature is constantly changing to reflect 

recent work, <strong>and</strong> some species are known by more than one name. Thus 

a complete listing of names, including synonyms, is an important starting 

point for locating information, whether as the basis for applied <strong>and</strong> basic 

research or simply to satisfy a curiosity. 

The information included in a catalog is usually arranged in a logical 

<strong>and</strong> organized format that allows for its convenient <strong>and</strong> rapid conveyance-in 

short, a quick <strong>and</strong> easy storage <strong>and</strong> retrieval system. The format <strong>and</strong> amount 

of information presented varies greatly, however. Our use of the term catalog 

is intended to convey a more comprehensive treatment, including information 

on all valid names, synonyms, type species, <strong>and</strong> the status <strong>and</strong> 

deposition of primary types. The bibliographic section includes complete 

references (author, date, original citation), <strong>and</strong> distributional <strong>and</strong> other biotic 

information are also provided. Not all citations that occur in the literature 

of Dryomyzidae are included in this catalog or the bibliographic section, 

especially where we suspect that the species being treated was misidentified, 

<strong>and</strong> inclusion would further promulgate inaccurate distributional data. 

Regional catalogs, checklists, <strong>and</strong> faunal treatments have contributed 

significantly to the compilation of this catalog. The most recent of these for 

a given region or country are as follows: 

<strong>Catalog</strong>s: Steyskal 1965b (Nearctic), 1977a (Neotropical), 1977b (Oriental); 

Sóos 1984 (Palearctic); Pitkin & Evenhuis 1989 (Australasian/Oceanian; 

see D. K. McAlpine 1995). 

Checklists: Hackman 1980 (Finl<strong>and</strong>); Martinek 1987 (Czech Republic, 

Slovakia); Morimoto 1989 (Japan); Nowakowski 1991 (Pol<strong>and</strong>); Gros-

WORLD CATALOG AND CONSPECTUS ON THE FAMILY DRYOMYZIDAE 209 

series 1991 (Belgium); Munari & Rivosecchi 1995 (Italy); Poole & Gentili 

1996 (Nearctic Region); Bächli 1998 (Switzerl<strong>and</strong>); Ch<strong>and</strong>ler 1998 (British 

Isles); Teschner 1999 (Germany); Pakalniškis et al. 2000 (Lithuania); Papp 

2001 (Pol<strong>and</strong>); Petersen & Meier 2001 (Denmark); Zuijlen & Beuk 2002 

(Netherl<strong>and</strong>s); Carles-Tolrá & Báez 2002 (Spain). 

Faunal Treatments: Czerny 1930 (Palearctic); Séguy 1934 (France); 

Kurahashi 1981 (Japan); Ozerov 1987 (Russia), 1998 (Palearctic), 1999 

(Far East, Russia); Bächli 1997 (Switzerl<strong>and</strong>) Falk 2005 (British Isles). 

Fossil: Hennig 1965, 1969; Evenhuis 1994. 

Acronyms used in this catalog 

To economize on space we have used acronyms for museums where primary 

type(s) are deposited. These acronyms are as follows: 

ABRI Azuma Biological Research Institute, Takarazuka, Hyogo, Japan. 

BLKU Biosystematics Laboratory, Graduate School of Social <strong>and</strong> Cultural 

Studies, Kyushu University, Fukuoka, Japan. 

CUI Cornell University, Ithaca, New York, USA. 

HUS Hokkaido University, Sapporo, Hokkaido, Japan. 

KLEBS Private collection of Prof. Dr. R. Klebs, Steinzeit. Konigsberg, 

Russia. 

LACM Los Angeles County Museum, Los Angeles, California, USA. 

MCZ Museum of Comparative Zoology, Harvard University, Cambridge, 

Massachusetts, USA. 

MNHNP Muséum National d’Histoire Naturelle, Paris, France. 

NMI National Museum of Irel<strong>and</strong>, Dublin, Irel<strong>and</strong>. 

NMW Naturhistorisches Museum, Wien, Austria. 

NSMT National Science Museum (Natural History), Tokyo, Japan. 

RNH Nationaal Natuurhistorische Museum, Leiden, Netherl<strong>and</strong>s. 

SPMC Sh<strong>and</strong>ong Provincial Museum, Jinan, China. 

USNM former United States National Museum, collection incorporated 

in the National Museum of Natural History, Smithsonian Institution, 

Washington, D.C., USA. 

ZIL Zoological Institute, Lund University, Lund, Sweden. 

ZISP Zoological Institute, Academy of Sciences, St. Petersburg, Russia. 

ZMHU Zoologisches Museum, Humboldt Universität, Berlin, Germany. 

ZMM Zoological Museum, Moscow University, Moscow, Russia.


Diagnosis. Adult. Medium-sized to large flies (body length 4.0-18.0 

mm); moderately robust, conspicuously setulose to moderately or strongly 

setose; yellow to brown or gray, moderately microtomentose. Head: 

Postocellar setae long, parallel or slightly divergent; 1-3 lateroclinate 

fronto-orbital setae. Antennae narrowly separated basally; antenna generally 

oriented ventrally; scape with setulae extended on to medial surface; 

pedicel short, lacking a complete dorsal seam. Face uniformly sclerotized, 

shallowly convex in profile; parafacial suture complete, extended with associated, 

shiny ridge to postgenal area; gena high (ca. 1/2-2/3 eye height), 

ventral half mostly setulose, dorsal half mostly bare; clypeus large, conspicuous, 

sometimes protruding. Thorax: Acrostichal setae either absent 

or with a prescutellar pair; dorsocentral setae 1-6; metepisternum bare; 

katepisternal suture more or less straight, not arched dorsally posteriorly; 

prosternum bare, straplike, extended laterally to join propleuron as a narrow, 

precoxal bridge; proepisternal seta long; proepimeral seta lacking. 

Wing hyaline to lightly infuscate; crossveins sometimes clouded; costa 

unbroken, bearing only setulae, not spinose; subcosta complete; apical 

section of vein M nearly straight, terminating posterior of wing apex; 

crossvein bm-cu distinctly apical of vein CuA 2 ; vein CuA 2 +A 1 evident 

to wing margin; vein A 2 beyond alula as a long, curved crease. Midtibia 

with posterior (variable), preapical, <strong>and</strong> apical setae only; basitarsi lacking 

terminal ventral processes; male forebasitarsus unmodified; apical 2 

tarsomeres flattened <strong>and</strong> somewhat exp<strong>and</strong>ed. Abdomen: Sternite 1 short 

but well sclerotized; sternites 1-2 incompletely separated; spiracles 2-5 in 

tergites. Female with 3 (1+2) small spermathecae. 

Egg. Creamy white; elongate (length 1.2-1.4 mm, width 0.4-0.5 mm), 

bearing a pair of flanges, extended dorsolaterally from sides; chorion with 

fine, honeycomb-like reticulation. 

Third-instar larva. Typically maggot-like, cylindrical, anterior end tapered, 

posterior end generally broadly rounded; anterior spiracle on short 

process, fan-shaped, digitate, with numerous terminal openings; posterior 

spiracles separate, on short or elongate respiratory tubes, each spiracle with 

3 slits, surrounded by a circlet of plumose hydrofuge hairs (not in fascicles); 

anal opening on protuberance, this sometimes subdivided into lobes 

that bear spines or star-like sclerites around anus. 

Puparium. Dark brown, ovoid with a moderately rugose surface. 

Adult Dryomyzidae are similar to Coelopidae <strong>and</strong> Helcomyzidae <strong>and</strong> 

are sometimes confused with Heleomyzidae <strong>and</strong> Scathophagidae. They 

are distinguished from these <strong>and</strong> other families of Schizophora, especially 

those in the superfamily Sciomyzoidea, by the following combination of


characters: postocellar setae long, parallel or slightly divergent; antennal 

bases approximate basally, narrowly separated; pedicel short, lacking a 

complete dorsal seam; oral vibrissae absent; face uniformly sclerotized, 

usually shallowly convex medially; clypeus large <strong>and</strong> prominent; greater 

ampulla lacking; prosternum bare, strap-like, well separated from propleuron; 

wing lacking costal spines; crossvein bm-cu aligned distinctly apical to 

vein CuA 2 ; some or all tibiae bearing a preapical dorsal seta. 

Biology. Two distinctly different biologies are demonstrated in the natural 

history of this family. The larvae of Oedoparena Curran are predators of intertidal 

barnacles (Burger et al. 1980, Suwa 1981) <strong>and</strong> are thus only known 

from maritime coasts. The other genera, where information is available, 

prefer humid environments such as forests where there is an under story of 

shaded, low vegetation <strong>and</strong> where decaying organic matter is found. Larvae 

in these habitats have been discovered to feed in rotting plants <strong>and</strong> fungi, 

dead animals ranging from insects <strong>and</strong> mollusks to vertebrate carrion <strong>and</strong> 

even in human excrement. The larvae are saprophagous, <strong>and</strong> females usually 

lay eggs directly on the rotting food source. 

Distribution. Species of Dryomyzidae are largely temperate in distribution 

<strong>and</strong> thus far are exclusively known from the northern hemisphere. With the 

exception of Oedoparena, which is strictly coastal in distribution, the other 

genera occur most frequently in humid, forested habitats. Oedoparena is 

only known from the cool, maritime coasts of the northern Pacific (Japan, 

Pacific Northwest of North America), <strong>and</strong> the other extant genera of the 

family are widespread across the Holarctic Region where their preferred 

habitats are found. Only four species occur in the Oriental Region, <strong>and</strong> 

these are found primarily in areas with temperate climates. As noted previously, 

the family is noticeably lacking in the Afrotropical, Australasian, <strong>and</strong> 

Neotropical Regions. 

Table of extant genera 

The following table lists the extant genera of Dryomyzidae in the order 

found in the catalog, with a summary of the number of extant species 

known from each zoogeographic region. The number in the total column is 

less than the sum of those in each region as species are known from more 

than one region.


Taxon 

NE NT 

DisTribuTion 

PA AF OR AU Total 

Dryomyza Fallén 2 8 2 10 

Dryope Robineau-Desvoidy 2 2 3 

oedoparena Curran 2 1 3 

Paradryomyza Ozerov 1 3 2 4 

Pseudoneuroctena Ozerov 1 1 1 

Steyskalomyza Kurahashi 1 1 

ToTals 8 16 4 22 

Classification <strong>and</strong> nomenclature. The concept of the family Dryomyzidae 

adopted here includes six extant <strong>and</strong> two fossil genera. The names of three 

genera (Dryomyza, Dryope, <strong>and</strong> Neuroctena) need clarification, as they 

have been confused, <strong>and</strong> errors concerning their usage have been perpetuated 

<strong>and</strong> promulgated in the literature. 

The type species of Dromyza has often been cited as D. vetula Fallén 

(= Musca flaveola Fabricius, 1794) through the subsequent designation of 

Westwood (1840: 165). Dryomyza vetula, however, was not an originally 

included species, <strong>and</strong> Westwood showed no synonymy that would link 

that species <strong>and</strong> name with any of the originally included species (Sabrosky 

1999). Thus, Westwood’s type designation is invalid. Zetterstedt 

(1846: 2082) published the next type designation for Dryomyza when he 

listed Dryomyza anilis Fallén as the type species. Dryomyza anilis is an 

originally included species, <strong>and</strong> Zetterstedt’s subsequent designation is 

thus valid. 

Robineau-Desvoidy (1830) proposed the genus Dryope for two of his 

species: D. communis <strong>and</strong> D. liturata. Coquillett (1910) subsequently designated 

Dryomyza communis (= Musca flaveola Fab., 1794) as the type 

species of Dryope, <strong>and</strong> as this species is an originally included species, this 

designation is valid <strong>and</strong> the generic name is available. 

The third generic name, Neuroctena Rondani, 1868, is invalid. The 

type species of Neuroctena by monotypy is Dryomyza anilis, which had 

previously been designated as the type species of Dryomyza (Zetterstedt 

1846). Neuroctena is thus an objective, junior synonym of Dryomyza <strong>and</strong> 

is an invalid name. 

Czerny (1930) <strong>and</strong> several subsequent authors (Séguy 1934, Hendel 

1937, Stackelberg 1970, Ozerov 1987, 1998, 1999, Ch<strong>and</strong>ler 1998, Zuijlen 

<strong>and</strong> Beuk 2002, Carles-Tolrá & Báez 2002) have recognized the genus 

“Neuroctena” as a separate genus. As noted above, however, Neuroctena 

is an invalid name, <strong>and</strong> the oldest available <strong>and</strong> valid name for Czerny


et al.’s generic concept of “Neuroctena” is Dryomyza. Czerny (1930) also 

recognized Dryomyza as a separate genus (separate from “Neuroctena”) 

<strong>and</strong> included D. flaveola <strong>and</strong> D. decrepita Zetterstedt in it. If D. flaveola 

<strong>and</strong> related species are recognized as belonging to a separate genus from 

Dryomyza, then the oldest available <strong>and</strong> valid name for this genus would 

be Dryope. In summary, the genus for which “Neuroctena” has sometimes 

been used in the literature (Czerny 1930, Séguy 1934, Hendel 1937, Stackelberg 

1970, Ozerov 1987, 1998, 1999, Ch<strong>and</strong>ler 1998, Zuijlen & Beuk 

2002, Carles-Tolrá & Báez 2002) is actually Dryomyza, <strong>and</strong> in these same 

cases, what has been called Dryomyza is Dryope. 

The phylogenetic relationships for the genera included in Dryomyzidae 

<strong>and</strong> for the families within the Sciomyzoidea have not been fully 

resolved (McAlpine 1991). Most phylogenetic considerations are either 

preliminary or they are concerned with the monophyly of a particular 

genus or small group of genera <strong>and</strong> not how these lineages are phylogenetically 

related to other such clades. J. F. McAlpine (1989), for example, 

suggested that the monophyly of the Dryomyzidae (Dryomyzinae in his 

sense) is supported by: (1) metasternum bare <strong>and</strong> (2) abdominal spiracles 

2-5 in the ventral margins of their respective tergites (except for Oedoparena 

minor Suwa <strong>and</strong> the fossil species Prodryomyza electrica Hennig). 

Mathis & Steyskal (1980) provided evidence on the monophyly of Oedoparena: 

(1) anepisternum setulose, (2) antennae approximate at base, 

(3) clypeus relatively well developed, (4) loss of acrostichal setae, <strong>and</strong> 

(5) an association with barnacles. However, the clypeal, acrostichal, <strong>and</strong> 

antennal characters occur elsewhere in the Dryomyzidae, <strong>and</strong> their phylogenetic 

importance is questionable <strong>and</strong> needs further assessment. The 

setose anepisternum <strong>and</strong> a close association with barnacles may indicate 

the monophyly of the genus. 

Although Ozerov (1998) divided the extant six genera into two tribes, 

Dryomyzini <strong>and</strong> Oedoparenini, that precedent is not followed here because 

the placement of the extinct genera within these tribes has not been clarified. 

Moreover, the tribes were not proposed within a phylogenetic context 

nor was evidence (synapomorphies) elaborated to support their recognition. 

Oedoparena, however, is certainly distinct in its morphology <strong>and</strong> natural 

history from other genera of the family <strong>and</strong> may merit recognition as a 

separate tribe. 

The genus Helcomyza Curtis <strong>and</strong> related genera have sometimes been 

classified as the subfamily Helcomyzinae within the Dryomyzidae (Czerny 

1930, Séguy 1934, Griffiths 1972, Steyskal 1987, McAlpine 1989). 

McAlpine (1991) enumerated several characters that distinguish the Helco-


myzidae as a family within the Sciomyzoidea <strong>and</strong> as separate from Dryomyzidae, 

Ropalomeridae, <strong>and</strong> Coelopidae. McAlpine further suggested the 

possibility that Helcomyzidae is the sister group to the Coelopidae. We follow 

McAlpine’s precedent <strong>and</strong> for the present, treat the Dryomyzidae, Helcomyzidae, 

Coelopidae, <strong>and</strong> Heterocheilidae as separate families. 

Key to extant genera of Dryomyzidae 

(modified from Ozerov 1998) 

1. Proepisternum <strong>and</strong> anepisternum densely setose, setae fine; clypeus 

tilted upward; prescutellar acrostichal setae absent; antennal 

bases closely approximate, distance between scapes narrower 

than width of anterior ocellus; dark bluish gray in ground color of 

body ................................................................ Oedoparena Curran 

- Proepisternum <strong>and</strong> anepisternum bare; clypeus not upturned; prescutellar 

acrostichal seta present (absent in Paradryomyza); antennal 

bases with distinct gap between, distance between scapes as 

long as or longer than width of anterior ocellus; yellow to brown 

in ground color of body ................................................................ 2 

2. Vein R 1 setose along length .................................Dryomyza Fallén 

- At least basal half or all of vein R 1 without distinct row of setae 3 

3. Postgena covered with hairs, <strong>and</strong> no setae ................................... 4 

- Postgena bearing many setae <strong>and</strong> occasionally bearing hairs ..... 5 

4. Arista bare. Postpronotum with 2 setae; katepisternum with 6 well 

developed setae dorsally in 2 rows. Dorsum of legs <strong>and</strong> abdomen 

mostly bearing well-developed setae ..... Steyskalomyza Kurahashi 

- Arista with short hairs. Postpronotum with 1 seta; katepisternum 

usually with 3 setae in almost 1 row. Dorsum of legs <strong>and</strong> abdomen 

mostly densely setulose, lacking well-developed setae .................. 

........................................................... Dryope Robineau-Desvoidy 

5. Prescutellar acrostichal seta absent. Hindfemur bearing 2 rows of 

spinules apically on ventral surface. Female abdominal tergite 7 

spinulose .....................................................Paradryomyza Ozerov 

- Prescutellar acrostichal seta present. Hindfemur <strong>and</strong> female abdominal 

tergite 7 lacking spinules ........Pseudoneuroctena Ozerov


Family Dryomyzidae Schiner 

Dryomyzinae Schiner, 1862a: 148 [as the subfamily Dryomyzinae, family 

Muscidae]. Type genus: Dryomyza Fallén, 1820b. 

Dryomyzidae. Brauer 1883: 39 [as a family]; Aldrich 1905: 578 [catalog, 

Nearctic]; Becker 1905: 37 [catalog, Palearctic]; Czerny 1930: 4 

[review, Palearctic]; Séguy 1934: 88 [France]; Crampton 1944: 153 

[classification]; Hennig 1958: 589 [discussion, phylogenetic relationships], 

1965: 73 [fossil fauna], 1973: 56 [discussion]; Stackelberg 

1970: 175 [key, Russia]; Hackman 1980: 144 [checklist, 

Finl<strong>and</strong>]; Kurahashi 1981: 437 [revision, Japan]; Barnes 1984: 43 

[biology, immature stages]; Sóos 1984: 153 [catalog, Palaearctic]; 

Martinek 1987: 177 [checklist, Czech Republic, Slovakia]; Ozerov 

1987: 40 [review, Palearctic], 1999: 555 [Russian Far East]; 

McAlpine 1989: 1450 [review, classification, phylogeny]; Morimoto 

1989: 805 [checklist, Japan]; Grosseries 1991: 133 [checklist, 

Belgium]; Nowakowski 1991: 180 [checklist, Pol<strong>and</strong>]; Munari & 

Rivosecchi 1995: 3 [checklist, Italy]; Poole & Gentili 1996: 153 

[checklist, Nearctic]; Bächli 1997: 30 [Switzerl<strong>and</strong>], 1998: 248 

[checklist, Switzerl<strong>and</strong>]; Ch<strong>and</strong>ler 1998: 131 [checklist, British 

Isles]; Teschner 1999: 148 [checklist, Germany]; Pakalniškis et al. 

2000: 33 [checklist, Lithuania]; Papp, 2001: 295 [checklist, Hungary]; 

Petersen & Meier 2001: 194 [checklist, Denmark]; Zuijlen 

& Beuk, 2002: 247 [catalog, Netherl<strong>and</strong>s]; Carles-Tolrá & Báez 

2002: 163 [checklist, Spain]. 

Dryomyzini. Schiner 1862a: 148 [as a tribe]. Ozerov 1998: 353 [Palaearctic 

manual]. 

Oedoparenini Ozerov, 1998: 353. Type genus: Oedoparena Curran, 1934. 

Genus DRYOMYzA Fallén 

Dryomyza Fallén, 1820a: 15. Type species: anilis Fallén, by subsequent 

designation [Zetterstedt 1846: 2082]. Zetterstedt 1846: 2082 [revision, 

Sc<strong>and</strong>inavia]; Schiner 1862b: 39 [Austria]; Becker 1905: 

37 [synonymy of Neuroctena]; Steyskal 1957: 56 [synonymy of 

Stenodryomyza]; Stackelberg 1970: 175 [key, Russia]; Hackman 

1980: 144 [checklist, Finl<strong>and</strong>]; Barnes 1984: 43 [biology, immature 

stages]; Sóos 1984: 153 [catalog, Palaearctic]; Martinek 

1987: 177 [checklist, Czech Republic, Slovakia]; Morimoto 1989: 

805 [checklist, Japan]; Grosseries 1991: 133 [checklist, Belgium];


Nowakowski 1991: 180 [checklist, Pol<strong>and</strong>]; Munari & Rivosecchi 

1995: 3 [checklist, Italy]; Poole & Gentili 1996: 153 [checklist, 

Nearctic]; Bächli 1997: 30 [Switzerl<strong>and</strong>], 1998: 248 [checklist, 

Switzerl<strong>and</strong>]; Sabrosky 1999: 119 [discussion; “The type species 

has often been cited as Musca flaveola Fabricius 1794 by 

designation of Westwood (1840: 145), but that nominal species 

was not originally included, <strong>and</strong> Westwood showed no synonymy 

that would link it with an originally included species.”]; Teschner 

1999: 148 [checklist, Germany]; Pakalniškis et al. 2000: 

33 [checklist, Lithuania]; Papp, 2001: 295 [checklist, Hungary]; 

Petersen & Meier 2001: 194 [checklist, Denmark]. 

Neuroctena Rondani, 1868: 56. Type species: Dryomyza anilis Fallén, by 

monotypy. Aldrich 1905: 578 [catalog, Nearctic]; Czerny 1930: 4 

[review, Palearctic]; Séguy 1934: 88 [France]; Hendel 1937: 186 

[generic key]; Stackelberg 1970: 175 [European Russia]; Kurahashi 

1981: 437 [revision, Japan]; Ozerov 1987: 40 [review, Palearctic], 

1998: 354 [Palearctic], 1999: 555 [Russian Far East]; 

Ch<strong>and</strong>ler 1998: 131 [checklist, British Isles]; Zuijlen & Beuk, 

2002: 247 [catalog, Netherl<strong>and</strong>s]; Carles-Tolrá & Báez 2002: 163 

[checklist, Spain]. Syn. Becker 1905. 

Neuroctena (Stenodryomyza) Hendel, 1924: 214. Type species: Scatophaga 

formosa Wiedemann, by original designation. Steyskal 1977b: 

173 [catalog, Orient]; Ozerov 1987: 431 [review, Russia], 1999: 

554 [Russia, Far East]. Syn. Steyskal 1957. 

Stenodryomyza. Hendel 1937: 186 [new status, generic key]. 

amblia Kurahashi. PA: Japan (Honshu). 

Dryomyza amblia Kurahashi, 1981: 443. Japan. Honshu: Ishikawa, Mt. 

Iozen. HT ♀ NSMT. 

anilis Fallén. NE: Canada (British Columbia, Labrador, Nova Scotia, Ontario, 

Quebec), USA (Alaska, Connecticut, Iowa, Illinois, Massachusetts, 

Michigan, Minnesota, New Hampshire, New Jersey, New 

York, North Carolina, Oregon, Pennsylvania, Rhode Isl<strong>and</strong>, South 

Dakota, Vermont, Washington). PA: Austria, Belgium, Czech Republic, 

Denmark, Finl<strong>and</strong>, France, Great Britain, Hungary, Italy, 

Japan (Honshu), Korea, Lithuania, Netherl<strong>and</strong>s, Pol<strong>and</strong>, Russia 

(European Part, Far East), Slovakia, Spain, Sweden, Switzerl<strong>and</strong>. 

Dryomyza anilis Fallén, 1820a: 16. Sweden. ST ♀♂ MNHNP [2♂, 

Becker 1902: 219], NMW, ZIL. Zetterstedt 1846: 2082 [revision, 

Sc<strong>and</strong>inavia]; Schiner 1862b: 39 [Austria]; Becker 1905: 37 [catalog, 

Palaearctic, synonymy of liturata], 1915: 128 [list]; Vimmer


1927: 40 [list]; Stackelberg 1970: 175 [key, Russia]; Mani 1974: 

222 [list]; Hackman 1980: 144 [checklist, Finl<strong>and</strong>]; Smith 1980: 167 

[larva, key]; Barnes 1984: 43 [biology, immature stages]; Nowakowski 

1991: 180 [checklist, Pol<strong>and</strong>]; Munari & Rivosecchi 1995: 

3 [checklist, Italy]; Poole & Gentili 1996: 153 [checklist, Nearctic]; 

Bächli 1997: 30 [Switzerl<strong>and</strong>], 1998: 248 [checklist, Switzerl<strong>and</strong>]; 

Papp, 2001: 295 [checklist, Hungary]; Petersen & Meier 2001: 194 

[checklist, Denmark]; Fast & Wheeler 2004: 11 [list, Quebec]. 

Dryope liturata Robineau-Desvoidy, 1830: 619. Not given (? France). 

ST ? MNHNP. Syn. Becker 1905. 

Neuroctena anilis. Rondani 1868: 56 [generic combination], 1869: 8 

[list]; Aldrich 1905: 578 [catalog, Nearctic synonymy of pallida]; 

Czerny 1930: 4 [review]; Séguy 1934: 88 [France]; Stackelberg 

1970: 175 [European Russia]; Ovchinnikova 1994: 3 [musculature 

of male terminalia]; Ch<strong>and</strong>ler 1998: 131 [checklist, British Isles]; 

Zuijlen & Beuk, 2002: 247 [catalog, Netherl<strong>and</strong>s]; Carles-Tolrá & 

Báez 2002: 163 [checklist, Spain]; Speight 2004: 49 [list, Irel<strong>and</strong>]; 

Falk 2005: 8 [key, Great Britain]; Papp 2005: 200 [list, Korea]. 

Dryomyza pallida Day, 1881: 89. USA. Connecticut. HT ? [apparently 

lost]. Syn. Aldrich 1905. 

Dryomyza melanacme Kurahashi, 1981: 441. Japan. Honshu: Ishikawa, 

Mt. Hakusan. HT ♀ NSMT. Syn. Ozerov 1999. 

Dryomyza analis. Misspelling. Sóos 1984: 153 [catalog, Palaearctic]; 

Martinek 1987: 177 [checklist, Czech Republic, Slovakia]; Grosseries 

1991: 133 [checklist, Belgium]; Teschner 1999: 148 [checklist, 

Germany]; Pakalniškis et al. 2000: 33 [checklist, Lithuania]. 

Neuroctena analis. Misspelling. Ozerov 1987: 40 [review, Russia, synonymy 

of melanacme], 1999: 555 [Russian Far East]. 

badia Kurahashi. PA: Japan (Honshu), Korea, Russia (Far East). 

Dryomyza badia Kurahashi, 1981: 440. Japan. Honshu: Nagano, Shimashima-dani. 

HT ♀ NSMT. Ozerov 1999: 555 [Russian Far East]. 

Neuroctena badia. Ozerov 1987: 40 [review, Russia, generic combination], 

1999: 555 [Russian Far East]; Papp 2005: 200 [list, Korea]. 

caucasica (Ozerov). PA: Russia (Caucasus). 

Neuroctena caucasica Ozerov, 1987: 40. Russia. Dagestan: Gunib. HT 

♀ ZISP. 

Dryomyza caucasica. NEW COMBINATION. 

ecalcarata Kurahashi. PA: Japan (Hokkaido, Honshu), Russia (Far East). 

Dryomyza ecalcarata Kurahashi, 1981: 442. Japan. Honshu: Toyama, 

Gokayama. HT ♀ NSMT.


Neuroctena ecalcarata. Ozerov 1987: 41 [review, Russia, generic 

combination], 1999: 554 [Russian Far East]. 

formosa (Wiedemann). OR: India, Taiwan, Vietnam. PA: China, Japan 

(Honshu, Kyushu, Shikoku), Korea, Russia (Far East). 

Scatophaga formosa Wiedemann, 1830: 447. Japan. ST ♀ ZMHU. 

Dryomyza maculipennis Macquart, 1851: 246. India. ST ♀ MNHNP. 

Osten Sacken 1882: 21 [synonymy]; Brunetti 1907: 169 [list, India]; 

Lichtwardt 1909: 127 [discussion, comparison with formosa]. 

Syn. Osten Sacken 1882. 

Dryomyza formosa. Loew 1858: 112 [generic combination]; Osten 

Sacken 1882: 20 [synonymy of maculipennis <strong>and</strong> gigas]; Wulp 

1896: 163 [south Asia]; Becker 1905: 37 [catalog, Palaearctic]; 

Matsumura 1905: 116 [review], 1931: 372 [review]; Shiraki 1932: 

47 [review]; Yasumatsu 1939: 419 [review]; Steyskal 1957: 63 

[revision]; Kurahashi 1981: 439 [revision]; Sóos 1984: 153 [catalog, 

Palaearctic]; Morimoto 1989: 805 [checklist, Japan]; McAlpine 

1995: 42 [new records, Fijian (isl<strong>and</strong> of Komo) - probably 

erroneous]. 

Dryomyza gigas Snellen von Vollenhoven, 1862: 18. Japan. ST ? RNH. 

Syn. osten Sacken 1882. 

Eggizoneura formosa. Coquillett 1898: 339 [generic combination]; Matsumura 

1905: 116 [review], 1931: 372 [review], 1932: 49 [review]. 

Neuroctena (Stenodryomyza) formosa. Hendel 1924: 214 [generic 

combination]; Wu 1940: 385 [China]; Steyskal 1977b: 173 [catalog, 

Orient]; Ozerov 1987: 431 [review, Russia], 1999: 554 [Russian 

Far East]; Papp 2005: 200 [list, Taiwan, Vietnam]. 

Stenodryomyza formosa. Czerny 1930: 5 [review, generic combination]; 

Hendel 1937: 186 [generic key]; Shiraki 1952: 1668 [review]; 

Takeuchi 1955: 151 [list]; Kanou 1959: 603 [larval morphology 

<strong>and</strong> biology]. 

pakistana Kurahashi. OR: Pakistan (Punjab). 

Dryomyza pakistana Kurahashi, 1989: 44. Pakistan. Punjab: Murree, 

2000 m. HT ♀ NSMT. 

pelidua Statz. PA: Germany (compression fossil). 

Dryomyza pelidua Statz, 1940: 146. Fossil; Germany. Rott: Siebengebirge 

(Oligocene). HT ? LACM. Evenhuis 1994: 429 [review]. 

puellaris Steyskal. PA: China. 

Dryomyza puellaris Steyskal, 1957: 65. China. Szechwan: Suifu. HT ♀ 

USNM. Sóos 1984: 153 [catalog, Palaearctic]. 

shanwangensis Zhang. PA: China (compression fossil).


Dryomyza shanwangensis Zhang, 1989: 364. Fossil; China. Sh<strong>and</strong>og: 

Shanwang, 22 km E Linqu (Miocene). HT ? SPMC. Evenhuis 

1994: 429 [review]. 

simplex Loew. NE: Canada (Ontario, Quebec), USA (Connecticut, Georgia, 

Massachusetts, Michigan, New Hampshire, New Jersey, New 

York, North Carolina, Pennsylvania, Tennessee, Vermont). 

Dryomyza simplex Loew, 1862: 128. USA. “Middle States.” ST ? MCZ 

[MCZ type number 13211]. Steyskal 1957: 67 [revision, symonymy 

of ruthae]; Poole & Gentili 1996: 153 [checklist, Nearctic]; 

Fast & Wheeler 2004: 11 [list, Quebec]. 

Neuroctena simplex. Hendel 1911: 370 [generic combination]. 

Neuroctena ruthae Brimley, 1925: 75. USA. North Carolina: Macon, 

Franklin. HT ♀ USNM [not found; Steyskal examined in 1957: 

67]. Syn. Steyskal 1957. 

takae Azuma. PA: Japan (Honshu). 

Dryomyza takae Azuma, 2001: 472. Japan. Honshu: Shimane, Daisencho, 

Masumizu Highl<strong>and</strong>. HT ♀ ABRI. 

Genus DRYOPE Robineau-Desvoidy 

Dryope Robineau-Desvoidy, 1830: 618. Type species: communis Robineau- 

Desvoidy [= Musca flaveola Fabricius], by subsequent designation 

[Coquillett 1910: 536]. 

Driope. Misspelling. Rondani 1868: 55 [list, Italy]. 

Dryomyza of authors, not Fallén. Czerny 1930: 3 [review]; Séguy 1934: 

87 [France]; Hendel 1937: 186 [generic key]; Stackelberg 1970: 

175 [key, European Russia]; Sóos 1984: 153 [catalog, Palaearctic]; 

Martinek 1987: 177 [checklist, Czech Republic, Slovakia]; 

Ozerov 1987: 37 [Russia], 1999: 553 [Russian Far East], 2000: 

357 [review, Nearctic]; Nowakowski 1991: 180 [Pol<strong>and</strong>]; Grosseries 

1991: 133 [checklist, Belgium]; Munari & Rivosecchi 

1995: 3 [checklist, Italy]; Bächli 1997: 31 [Switzerl<strong>and</strong>], 1998: 

248 [checklist, Switzerl<strong>and</strong>]; Ch<strong>and</strong>ler 1998: 131 [checklist, British 

Isles]; Teschner 1999: 148 [checklist, Germany]; Pakalniškis 

et al. 2000: 33 [checklist, Lithuania]; Papp 2001: 295 [checklist, 

Hungary]; Petersen & Meier 2001: 194 [checklist, Denmark]; 

Carles-Tolrá & Báez 2002: 163 [checklist, Spain]; Zuijlen & Beuk 

2002: 247 [catalog, Netherl<strong>and</strong>s]. 

decrepita (Zetterstedt). NE: Canada (British Columbia, Nova Scotia), USA 

(Alaska, Kansas, Massachusetts, Michigan, New Hampshire, New


York, Pennsylvania, Vermont). PA: Austria, Belgium, Czech Republic, 

Germany, Great Britain, Irel<strong>and</strong>, Netherl<strong>and</strong>s, Pol<strong>and</strong>, Slovakia, 

Sweden, Switzerl<strong>and</strong>, Russia (European Part, Siberia, Far East). 

Dryomyza decrepita Zetterstedt, 1838: 737. Sc<strong>and</strong>inavia (various localities). 

ST ♀♂ ZIL. Zetterstedt 1846: 2085 [revision, Sc<strong>and</strong>inavia]; 

Schiner 1862b: 40 [Europe]; Becker 1905: 37 [catalog, Palaearctic]; 

Czerny 1930: 3 [review]; Steyskal 1957: 61 [as possible synonym 

of flaveola], 1962: 71 [discussion, included as synonym of flaveola]; 

Stackelberg 1970: 175 [key, European Russia]; Sóos 1984: 

153 [catalog, Palaearctic]; Martinek 1987: 177 [checklist, Czech 

Republic, Slovakia]; Nowakowski 1991: 180 [Pol<strong>and</strong>]; Grosseries 

1991: 133 [checklist, Belgium]; Bächli 1997: 31 [Switzerl<strong>and</strong>], 

1998: 248 [checklist, Switzerl<strong>and</strong>]; Ch<strong>and</strong>ler 1998: 131 [checklist, 

British Isles]; Ozerov 1999: 553 [Russian Far East], 2000: 357 

[review, synonymy of ferruginea <strong>and</strong> dayi]; Teschner 1999: 148 

[checklist, Germany]; Zuijlen & Beuk 2002: 247 [catalog, Netherl<strong>and</strong>s]; 

Falk 2005: 8 [key, discussion, Great Britain, Irel<strong>and</strong>]. 

Dryope decrepita. NEW COMBINATION. 

Dryomyza dayi Cresson, 1920: 34. USA. New York: Tomkins, Ithaca. 

HT ♀ CUI. Steyskal 1957: 62 [as synonym of D. flaveola]. Syn. 

Ozerov 2000. 

Dryomyza ferruginea Mel<strong>and</strong>er, 1920: 311. USA. Vermont: Caledonia, 

Lyndon. LT ♀ USNM [designated by Ozerov 2000: 357]. Poole & 

Gentili 1996: 153 [checklist, Nearctic]. Syn. Ozerov 2000. 

Dryomyza flaveola of Steyskal [not Fabricius]. Misidentification. 

Steyskal 1957: 61 [synonymy, discussion], 1958: 138 [new records 

(Alaska, British Columbia, Manitoba, Ontarion, Quebec, 

Nova Scotia)], 1962: 71 [as senior synonym of decrepita]. 

flaveola (Fabricius). PA: Austria, Belarus, Belgium, Czech Republic, Denmark, 

France, Germany, Great Britain, Hungary, Irel<strong>and</strong>, Italy, 

Lithuania, Netherl<strong>and</strong>s, Pol<strong>and</strong>, Slovakia, Sweden, Switzerl<strong>and</strong>, 

Russia (European Part, Caucasus), Ukraine. 

Musca flaveola Fabricius, 1794: 343. Denmark. Copenhagen. LT ♂ 

MNHNP [designated by Becker 1902: 219]. 

Dryomyza flaveola. Meigen 1826: 256 [generic combination]; Zetterstedt 

1846: 2084 [revision, Sc<strong>and</strong>inavia]; Schiner 1862b: 40 [Austria]; 

Becker 1902: 219 [review of Meigen’s types, synonymy of 

fuscicornis], 1905: 37 [catalog, Palaearctic, synonymy of communis, 

mollis, zawadskii <strong>and</strong> vetula]; Czerny 1930: 3 [review]; Séguy 

1934: 88 [France]; Hendel 1937: 186 [generic key]; Stackelberg


1970: 175 [key, European Russia]; Hackman 1980: 144 [checklist, 

Finl<strong>and</strong>]; Sóos 1984: 153 [catalog, Palaearctic]; Martinek 1987: 

177 [checklist, Czech Republic, Slovakia]; Nowakowski 1991: 

180 [Pol<strong>and</strong>]; Grosseries 1991: 133 [checklist, Belgium]; Munari 

& Rivosecchi 1995: 3 [checklist, Italy]; Bächli 1997:31 [Switzerl<strong>and</strong>], 

1998: 248 [checklist, Switzerl<strong>and</strong>]; Ch<strong>and</strong>ler 1998: 131 

[checklist, British Isles]; Teschner 1999: 148 [checklist, Germany]; 

Pakalniškis et al. 2000: 33 [checklist, Lithuania]; Papp 2001: 

295 [checklist, Hungary]; Petersen & Meier 2001: 194 [checklist, 

Denmark]; Zuijlen & Beuk 2002: 247 [catalog, Netherl<strong>and</strong>s]; Carles-Tolrá 

& Báez 2002: 163 [checklist, Spain]; Falk 2005: 8 [key, 

Great Britain]. 

Dryope flaveola. NEW COMBINATION. 

Dryomyza vetula Fallén, 1820a: 16. Sweden. “Beckaskog, Esperöd.” 

ST ♀♂ ZIL. Syn. Becker 1905. 

Dryope communis Robineau-Desvoidy, 1830: 619. Not given (? 

France). ST ? MNHNP. Syn. Becker 1905. 

Dryomyza mollis Haliday, 1833: 167. United Kingdom. Northern Irel<strong>and</strong>: 

Arde, Holywood. ST ? NMI. Syn. Becker 1905. 

Dryomyza zawadskii Schummel, 1834: 740. Pol<strong>and</strong>. “Silesia.” ST ? 

DEI. Schiner 1862b: 40 [Austria]; Hensel 1870: 133 [review]. 

Syn. Becker 1905. 

Dryomyza fuscicornis Meigen, 1838: 343. Germany. Bavaria. LT ♀ 

MNHNP [designated by Becker 1902: 219]. Schiner 1862b: 40 

[Germany]. Syn. Becker 1902. 

Dryomyza fusciformis. Misspelling. Sóos 1984: 153. 

mel<strong>and</strong>eri (Steyskal). NE: Canada (British Columbia, Quebec), USA 

(Alaska, Idaho). 

Dryomyza maculipennis Mel<strong>and</strong>er, 1920: 311. USA. Idaho: Latah, 

Moscow Mountain. LT ♀ USNM [designated by Ozerov 2000: 

358]. Preoccupied Macquart 1851. 

Dryomyza mel<strong>and</strong>eri Steyskal, 1957: 63 [new name for maculipennis 

Mel<strong>and</strong>er]. Poole & Gentili 1996: 153 [checklist, Nearctic]; Ozerov 

2000: 358 [review]. 

Dryope mel<strong>and</strong>eri. NEW COMBINATION. 

Genus OEDOPARENA Curran 

Oedoparena Curran, 1934: 382. Type species: glauca Coquillett, by original 

designation. Steyskal 1958: 141 [review], 1965b: 681 [catalog,


Nearctic]; Mathis & Steyskal 1980: 349 [revision]; Burger et al. 

1980: 360 [natural history]; Suwa 1981: 29 [revision]; Soós 1984: 

154 [catalog, Palaearctic]; Morimoto 1989: 805 [checklist, Japan]. 

glauca (Coquillett). NE: Canada (British Columbia), USA (Alaska, California, 

Washington). 

Oedoparea glauca Coquillett, 1900: 458. USA. Alaska: Metlakahtla. 

LT ♀ USNM [USNM type number 5254] [designated by Steyskal 

1958: 142]. 

Heterocheila glauca. Aldrich 1905: 578 [generic combination]. 

Heteromyza glauca. Mel<strong>and</strong>er 1920: 309 [generic combination]. 

Oedoparena glauca. Curran 1934: 382 [generic combination]; Cresson 

1920: 31 [review]; Saunders 1928: 552 [list]; Steyskal 1958: 141 

[review, figs. of male terminalia], 1965b: 681 [catalog, Nearctic]; 

Mathis & Steyskal 1980: 352 [revision]; Burger et al. 1980: 360 

[natural history]; Suwa 1981: 30 [review, figs. of male terminalia]. 

minor Suwa. PA: Japan (Hokkaido). 

Oedoparena minor Suwa, 1981: 30. Japan. Hokkaido: Otarushi, Asari 

beach. HT ♀ HUS. Soós 1984: 154 [catalog, Palaearctic]; Morimoto 

1989: 805 [checklist, Japan]. 

nigrifrons Mathis & Steyskal. NE: USA (Oregon, Washington). 

Oedoparena nigrifrons Mathis & Steyskal, 1980: 357. USA. Washington: 

Pacific, Ilwaco. HT ♀ USNM [USNM type number 76446]. 

Genus PALAEOTIMIA Meunier 

Palaeotimia Meunier, 1908: 266. Type species: ihoesti Meunier, by monotypy. 

ihoesti Meunier. PA: Baltic Region (amber). 

Palaeotimia ihoesti Meunier, 1908: 266. Fossil; Baltic Region. HT ♂ 

KLEBS [collection number 2395]. Evenhuis 1994: 429 [review]. 

Genus PARADRYOMYzA Ozerov 

Paradryomyza Ozerov, 1987: 38. Type species: Odontomera setosa Bigot, 

by original designation. 

orientalis Ozerov & Sueyoshi. OR: Taiwan. 

Paradryomyza orientalis Ozerov & Sueyoshi, 2002: 564. Taiwan. Taichun 

xian: Hoping xiang, Suchilanchi. HT ♀ BLKU. Papp 2005: 

200 [list, Taiwan].


setosa (Bigot). NE: Canada (British Columbia, Quebec), USA (Alaska, 

Colorado, Idaho, Massachusetts, Montana, New Hampshire, Vermont, 

Washington). PA: Russia (Siberia, Far East). 

Odontomera ? setosa Bigot, 1886: 386. USA. Washington. HT ♀ 

MNHNP. Hendel 1911: 370 [discussion]. 

Neuroctena setosa. Hendel 1911: 370 [generic combination, discussion, 

synonymy of fumida]. 

Dryomyza setosa. Steyskal 1957: 66 [generic combination]; Poole & 

Gentili 1996: 153 [checklist, Nearctic]. 

Paradryomyza setosa. Ozerov 1987: 38 [review, generic combination], 

1999: 553 [Russian Far East]. 

Neuroctena fumida Coquillett, 1901: 616. USA. New Mexico: Rio Arriba, 

Beulah. HT ♀ USNM [USNM type number 5504]. Aldrich 

1905: 579 [catalog, Nearctic]. Syn. Hendel 1911. 

spinigera Ozerov. PA: Russia (Far East). 

Paradryomyza spinigera Ozerov, 1987: 39. Russia. Far East: Amur Territory, 

Zeya. HT ♀ ZMM. Ozerov 1999: 554 [Russian Far East]. 

steyskali Ozerov & Sueyoshi. OR: India (Uttar Pradesh), Nepal. 

Paradryomyza steyskali Ozerov & Sueyoshi, 2002: 567. Nepal. 

Dobang Kharka: 83°24’E, 28°36’N, 2400 m. HT ♀ BLKU. 

Genus PRODRYOMYzA Hennig 

Prodryomyza Hennig, 1965: 73. Type species: electrica Hennig, by monotypy. 

electrica Hennig. PA: Baltic Region (amber). 

Prodryomyza electrica Hennig, 1965: 73. Fossil; Baltic Region. HT ♀ 

Institut der Universität Tübingen [type number 1299/3]. Evenhuis 

1994: 429 [review]. 

Genus PSEuDONEuROCTENA Ozerov 

Pseudoneuroctena Ozerov, 1987: 41. Type species: Dryomyza senilis 

Zetterstedt, by original designation. 

senilis (Zetterstedt). NE: USA (Alaska). PA: Finl<strong>and</strong>, Korea, Sweden, Russia 

(Siberia). 

Dryomyza flaveola of Zetterstedt [not Fabricius]. Misidentification. 

Zetterstedt 1838: 736 [list, Sweden]. 

Dryomyza senilis Zetterstedt, 1846: 2087. “Scilicet in Jamtl<strong>and</strong>iae 

paroecia Undersåker passim, ex. gr. ad Ristansfors, Åreskutan 

& Mullfjellen; in Lapponia Umensi ad Wilhelmina.” ST ? ZIL.


Becker 1905: 37 [catalog, Palaearctic]; Czerny 1930: 4 [review]; 

Séguy 1934: 88 [key]; Hackman 1980: 144 [checklist, Finl<strong>and</strong>]; 

Sóos 1984: 154 [catalog, Palaearctic]. 

Pseudoneuroctena senilis. Ozerov 1987: 41 [review, Russia, generic 

combination, synonymy of bergi]; Papp 2005: 200 [list, Korea]. 

Dryomyza bergi Steyskal, 1957: 60. USA. Alaska: Lower Yukon River. 

HT ♀ USNM. Kurahashi 1981: 438 [review]; Poole & Gentili 

1996: 153 [checklist, Nearctic]. Syn. Ozerov 1987. 

Genus STEYSKALOMYzA Kurahashi 

Steyskalomyza Kurahashi, 1982: 36. Type species: hasegawai Kurahashi, 

by original designation. Ozerov 1987. 

hasegawai Kurahashi. PA: Japan (Honshu). 

Steyskalomyza hasegawai Kurahashi, 1982: 37. Japan. Honshu: Iwate, 

Morioka, Kuriyagawa. HT ♀ NSMT. Iwasa 2002: 133 [biology]. 

Nomina dubia 

dubia Macquart. NT: “Patrie inconnue.” 

Dryomyza dubia Macquart, 1844: 343. “Patrie inconnue.” HT ♀ 

MNHNP. 

maculipes Walker. NT: Mexico. 

Dryomyza maculipes Walker, 1861: 319. Mexico. ST ? BMNH. 

Genus <strong>and</strong> species transferred to other families 

Genus SCIOGRIPHONEuRA Malloch 

Sciogriphoneura Malloch, 1933: 300. Type species: nigriventris Malloch, 

by original designation. Steyskal 1977: 1 [catalog, Neotropics]. 

Barnes 1981: 64-66 [transferred Sciogriphoneura <strong>and</strong> its two included 

species from Dryomyzidae to Helosciomyzidae]. 

brunnea Steyskal. NT: Brazil. 

Sciogriphoneura brunnea Steyskal, 1977: 1. Brazil. Rio de Janeiro: 

Itatiaia (Fazenda Serra). HT ♀ MZSP. 

nigriventris Malloch. NT: Chile. 

Sciogriphoneura nigriventris Malloch, 1933: 301. Chile. Chiloé: Ancud. 

HT ♀ BMNH. Steyskal 1977: 1 [catalog, Neotropics].


advena Wiedemann. NT: “South America.” 

Dryomyza advena Wiedemann, 1830: 449 [Helosciomyzidae, genus 

Sciosapromyza]. ST ♂ NHMW. Hendel 1933: 70 [transferred advena 

Wiedemann to Sciosapromyza Hendel as the type species of 

that genus by monotypy]. 

semicyanea Walker. Au: Indonesia (Maluku), PNG. 

Dryomyza semicyanea Walker, 1858: 109. Indonesia. Maluku. ST ? 

BMNH. 

Lamprogaster semicyanea. McAlpine 1995: 42 [generic combination, 

Platystomatidae]. 

Acknowledgments 

We gratefully thank <strong>and</strong> acknowledge the following people <strong>and</strong> their supporting 

institutions for assistance: F. C. Thompson (Systematic Entomology 

Laboratory, USDA, c/o Smithsonian Institution, Washington, D.C. 

USA) for advice on nomenclatural matters <strong>and</strong> literature; I. Brake (The 

Natural History Museum, London, UK) for general reviews <strong>and</strong> literature, 

S. Gaimari (California Food <strong>and</strong> Agriculture, Sacramento) for clarifying 

the status of Dryomyza advena Wiedemann <strong>and</strong> David K. McAlpine (Australian 

Museum, Sydney, Australia) for his review. 

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