Classification; pp. 21 - Nearctica

NOMINA INSECTA NEARCTICA 

THE CLASSIFICATION USED IN THIS VOLUME 

The classification of the Coleoptera used in this volume follows the following reference: 

Lawrence, J.F., and A. F. Newton. Families and subfamilies of Coleoptera (with selected genera, notes, 

references and data on family-group names), pp. 779-1006. in Pakaluk, J., and S.A. Slipinski. 1995. 

Biology, Phylogeny, and Classification of Coleoptera. Volume 2. Warsaw. Muzeum I Instytut Zoologii 

PAN. in two volumes, 1092 pp. This classification represents substantial changes from more traditional 

classifications, and a few family names may be somewhat overly split. However this reference will 

probably serve as the standard for the order Coleoptera for some time to come. The single exception is the 

retention of the groups Scolytidae and Platypodidae as separate families rather than subfamilies of thte 

Curculionidae. Some controversy seems to still exist about the status of these two groups. The family 

status of each has been retained because of economic importance, distinctiveness, and size of the 

Scolytidae. The family names are arranged as in Lawrence and Newton (1995) followed by the subfamilies 

applicable to the North American fauna (north of Mexico). The North American genera falling within each 

subfamily are listed alphabetically under the subfamily name. Generic names whose placement is not 

known to the editors are placed in a category in the family called unplaced. The term "unplaced" only 

denotes that the name is not associated with a subfamily assignment in the database used in assembling this 

list. The genus group name may be, and commonly is, assigned in the original description or a subsequent 

reference. The substantial list of references in the book and paper cited above may be consulted for more 

detailed phylogenies. The reference Arnett, R. H. 1971. Beetles of the United States (A Manual for 

Identification). Ann Arbor. American Entomological Institute. 1112 pp., is also extremely useful. A 

more detailed account of the current classification does not fall within the purpose of this list. 

SUBORDER ARCHOSTEMATA KOLBE 1908 

MICROMALTHIDAE Barber 1913 

Micromalthus. 

CUPEDIDAE Laporte 1836 

Cupes, Priacma. 

SUBORDER MYXOPHAGA CROWSON 1955 

HYDROSCAPHIDAE Leconte 1874 

Hydroscapha. 

MICROSPORIDAE Crotch 1873 

Microsporus. 

SUBORDER ADEPHAGA SCHELLENBERG 1806 

GYRINIDAE Latreille 1810 

Spanglerogyrinae Folkerts 1979: Spanglerogyrus. 

Gyrininae Latreille 1810: Dineutus, Gyretes, Gyrinus. 

HALIPLIDAE Aube 1836 

Apteraliplus, Haliplus, Peltodytes. 

TRACHYPACHIDAE Thomson 1857 

Trachypachus. 

NOTERIDAE Thomson 1860 

Noterinae Thomson 1860: Hydrocanthus, Notomicrus, Suphis, Suphisellus. 

AMPHIZOIDAE Leconte 1853 

21 

CLASSIFICATION

22 

CLASSIFICATION 


Amphizoa. 

DYTISCIDAE Leach 1815 

Copelatinae Van den Branden 1885: Agaporomorphus, Copelatus. 

Laccophilinae Gistel 1856: Laccodytes, Laccophilus. 

Hydroporinae Aube 1836: Andocheilus, Bidessonotus, Bidessus, Brachyvatus, Celina, 

Deronectes, Derovatellus, Desmopachria, Hydroporus, Hydrovatus, Hygrotus, Laccornis, 

Liodessus, Lioporeus, Neobidessus, Neoclypeodytes, Oreodytes, Pachydrus, Potamonectes. 

Colymbetinae Erichson 1837: Agabetes, Agabinus, Agabus, Carrhydrus, Colymbetes, 

Coptotomus, Hoperius, Hydrotrupes, Ilybius, Matus, Neoscutopterus, Rhantus. 

Dytiscinae Leach 1815: Acilius, Cybister, Dytiscus, Eretes, Graphoderus, Hydaticus, Megadytes, 

Thermonetus. 

RHYSODIDAE Laporte 1840 

Clinidium, Omoglymmius. 

CARABIDAE Latreille 1802 

Paussinae Latreille 1807: Metrius, Ozaena, Pachyteles, Physea. 

Gehringiinae Darlington 1933: Gehringia. 

Omophroninae Bonelli 1810: Omophron. 

Carabinae Latreille 1802: Calosoma, Carabus, Cychrus, Leistus, Nebria, Notiophilus, Opisthius, 

Pelophila, Scaphinotus, Sphaeroderus. 

Cicindelinae Latreille 1802: Amblycheila, Cicindela, Megacephala, Omus. 

Loricerinae Bonelli 1810: Loricera. 

Elaphrinae Latreille 1802: Blethisa, Diacheila, Elaphrus. 

Scaritinae Bonelli 1810: Ardistomis, Aspidoglossa, Clivina, Dyschirius, Halocoryza, 

Oxydrepanus, Pasimachus, Promecognathus, Scarites, Schizogenius, Semiardistomis. 

Trechinae Bonelli 1810: Ameroduvalius, Anillaspis, Anillinus, Anillodes, Asaphidion, 

Bembidion, Blemus, Broscodera, Broscus, Darlingtonea, Diplochaetus, Diplous, Elaphropus, 

Horologion, Micranillodes, Micratopus, Mioptachys, Miscodera, Neaphaenops, Nelsonites, 

Nomius, Paratachys, Patrobus, Pericompsus, Phrypeus, Platidiolus, Platypatrobus, Pogonistes, 

Pogonus, Polyderis, Porotachys, Pseudanophthalmus, Psydrus, Tachys, Tachyta, 

Thalassotrechus, Trechoblemus, Trechus, Troglanillus, Xenotrechus, Zacotus. 

Harpalinae Bonelli 1810: Abaris, Abax, Acupalpus, Agonum, Agra, Amara, Amblygnathus, 

Amerinus, Amphasia, Anatrichis, Anchomenus, Anchonoderus, Anisodactylus, Apenes, Apristus, 

Atranus, Axinopalpus, Aztecarpalus, Badister, Bradycellus, Calathus, Calleida, Calybe, 

Chlaenius, Colliuris, Colpodes, Coptodera, Cratacanthus, Cyclotrachelus, Cylindronotum, 

Cymindis, Dercylinus, Dicaelus, Dicheirus, Diplocheila, Discoderus, Dromius, Elliptoleus, 

Eucaerus, Euphorticus, Euproctinus, Euryderus, Evolenes, Galerita, Gastrellarius, Geopinus, 

Harpalus, Hartonymus, Helluomorphoides, Infernophilus, Inna, Lachnocrepis, Lachnophorus, 

Laemostenus, Lebia, Leptotrachellus, Licinus, Lophoglossus, Loxandrus, Micrixys, Microlestes, 

Morion, Myas, Nemotarsus, Notiobia, Olisthopus, Onota, Oodes, Oodinus, Ophryogaster, 

Oxycrepis, Oxypselaphus, Panagaeus, Paranchus, Pelmatellus, Pentagonica, Perigona, 

Philorhizus, Phloeoxena, Piesmus, Piosoma, Platynus, Plochionus, Poecilus, Pogonodaptus, 

Polpochila, Pseudamara, Pseudaptinus, Pterostichus, Rhadine, Selenophorus, Sericoda, 

Somotrichus, Stenocrepis, Stenolophus, Stenomorphus, Stereocerus, Stomis, Syntomus, Synuchus, 

Tanystoma, Tecnophilus, Tetragonoderus, Tetraleucus, Thalpius, Trichocellus, Trichotichnus, 

Xestonotus, Zuphium. 

Pseudomorphinae Newman 1842: Pseudomorpha. 

Brachininae Bonelli 1810: Brachinus, 

SUBORDER POLYPHAGA EMERY 1886 

Superfamily HYDROPHILOIDEA Latreille 1802


HYDROPHILIDAE Latreille 1802 

Helophorinae Leach 1815: Helophorus. 

Epimetopinae Zaitzev 1908: Epimetopus. 

Georissinae Laporte 1840: Georissus. 

Hydrochinae Thomson 1859: Hydrochus. 

Spercheinae Erichson 1837: Spercheus. 

Hydrophilinae Latreille 1802: Ametor, Anacaena, Berosus, Crenitis, Cymbiodyta, Derallus, 

Dibolocelus, Enochrus, Helobata, Helochares, Helocombus, Hydrobiomorpha, Hydrobius, 

Hydrochara, Hydrophilus, Laccobius, Paracymus, Sperchopsis, Tropisternus. 

Sphaeridiinae Latreille 1802: Agna, Cercyon, Cryptopleurum, Dactylosternum, Genyon, 

Ictenosternum, Megasternum, Omicrus, Oosternum, Pelosoma, Pemelus, Phaenonotum, 

Sphaeridium. 

SPHAERITIDAE Shuckard 1839 

Sphaerites. 

HISTERIDAE Gyllenhal 1808 

Abraeinae MacLeay 1819: Abraeus, Acritus, Aeletes, Halacritus, Plegaderus, Teretriosoma, 

Teretrius. 

Saprininae Blanchard 1845: Aphelosternus, Chelyoxenus, Eremosaprinus, Euspilotus, 

Geomysaprinus, Gnathoncus, Hypocaccus, Neopachylopus, Saprinus, Xerosaprinus. 

Dendrophilinae Reitter 1909: Anapleus, Bacanius, Carcinops, Dendrophilus, Geocolus, 

Paromalus, Platylomalus, Xestipyge. 

Onthophilinae MacLeay 1819: Onthophilus. 

Tribalinae Bickhardt 1914: Caerosternus, Epierus, Peploglyptus, Pseudepierus, Stictostix. 

Histerinae Gyllenhal 1808: Atholus, Baconia, Hister, Hololepta, Iliotona, Margarinotus, 

Merohister, Omalodes, Operclipygus, Phelister, Platysoma, Psiloscelis, Spilodiscus, Tribalister. 

Hetaeriinae Marseul 1857: Hetaerius, Mroczkowskiella, Reninus, Terapus, Ulkeus, Yarmister. 

Superfamily STAPHYLINOIDEA Latreille 1802 

HYDRAENIDAE Mulsant 1844 

Hydraeninae Mulsant 1844: Hydraena, Limnebius. 

Ochthebiinae Thomson 1859: Gymnochthebius, Neochthebius, Ochthebius. 

PTILIDAE Erichson 1845 

Ptiliinae Erichson 1845: Actidium, Dilinium, Macdonaldium, Micridium, Motschulskium, 

Nossidium, Ptenidium, Pteryx, Ptiliola, Ptiliolum, Ptilium. 

Nanosellinae Barber 1924: Bambara, Eurgyne, Hydnosella, Nanosella, Porophila, Suturina, 

Throscoptilium. 

Cephaloplectinae Sharp 1883: Limulodes, Paralimuodes. 

Acrotrichinae Reitter 1909: Acrotrichis, Actinopteryx, Nephanes, Ptiliodina, Ptiliopycna, 

Ptinella, Ptinellodes, Smicrus. 

AGYRTIDAE Thomson 1859 

Agyrtes, Apteroloma, Ipelates, Lyrosoma, Necrophilus, Pteroloma. 

LEIODIDAE Fleming 1821 

Catopocerinae Hatch 1927: Catopocerus, Glacicavicola. 

Leiodinae Fleming 1821: Agathidium, Aglyptinus, Anisotoma, Anogdus, Apheloplastus, 

Caenocyrta, Cainosternum, Colenis, Cyrtusa, Ecarinosphaerula, Hydnobius, Isoplastus, Leiodes, 

Lionothus, Neocyrtusa, Stetholioides, Triarthron. 

Coloninae Horn 1880: Colon. 

Cholevinae Kirby 1837: Adelopsis, Caloptrichus, Catops, Dissochaetus, Echinocoleus, Nemadus, 

Platycholeus, Prionochaeta, Ptomaphagus, Sciodrepoides. 

Platypsyllinae Ritsema 1869: Leptinillus, Leptinus, Platypsyllus. 

23 

CLASSIFICATION

24 



SCYDMAENIDAE Leach 1814 

Mastiginae Fleming 1821: Papsus. 

Scydmaeninae Leach 1815: Acholerops, Ascydmus, Catalinus, Cephennium, Ceramphis, 

Chevrolatia, Connophron, Delius, Drastophus, Euconnus, Eumicrus, Eutheia, Euthiodes, 

Leptoscydmus, Lophioderus, Neladius, Noctophus, Opresus, Pycnophus, Scydmaenus, Veraphis. 

SILPHIDAE Latreille 1807 

Silphinae Latreille 1807: Aclypea, Heterosilpha, Necrodes, Necrophila, Oiceoptoma, Oxelytrum, 

Thanatophilus. 

Nicrophorinae Kirby 1837: Nicrophorus. 

STAPHYLINIDAE Latreille 1802 

Omalinae MacLeay 1825: Acidota, Acrulia, Amphicroum, Anotylus, Anthobium, Arpedium, 

Artochia, Boreaphilus, Brathinus, Coryphium, Elonium, Ephelinus, Eucnecosum, Eudectoides, 

Eusphalerum, Gnathoryphium, Haida, Hapalaraea, Holoboreaphilus, Lesteva, Mathrilaeum, 

Micralymma, Microedus, Olophrum, Omalium, Omalonomus, Omalorphanus, Orobanus, 

Orochares, Pelecomalium, Phlaeopterus, Phloeonomus, Porrhodites, Psephidonus, Pycnoglypta, 

Subhaida, Tanyrhinus, Trigonodemus, Unamis, Vellica, Xenicopoda, Xylodromus. 

Empelinae Newton and Thayer 1992: Empelus 

Proteininae Erichson 1839: Megarthrus, Proteinus. 

Micropeplinae Leach 1815: Kalissus, Micropeplus 

Dasycerinae Reitter 1887: Dasycerus. 

Pselaphinae Latreille 1802: Abdiunguis, Acolonia, Actiastes, Actium, Actizona, Adranes, 

Allobrox, Allotrimium, Anchylarthron, Anitra, Arianops, Arthmius, Atinus, Basolum, 

Batriasymmodes, Batrisodes, Bibloplectus, Bibloporus, Biotus, Bontomtes, Brachygluta, 

Briaraxis, Bythinoplectus, Bythinopsis, Caccoplectus, Cedius, Ceophyllus, Cercocerus, 

Conoplectus, Ctenisis, Cupila, Custotychus, Cylindrarctus, Dalmodes, Dalmosella, Decarthron, 

Euboarhexius, Euphalepus, Euplecturga, Euplectus, Eupsenius, Eutrichites, Eutyphlus, 

Foveoscapha, Fustiger, Hamotus, Hatchia, Hesperotychus, Lemelba, Leptoplectus, Lucifotychus, 

Machaerites, Machaerodes, Mayetia, Megarafonus, Melba, Mipselytrus, Morius, Nearctitychus, 

Nisaxis, Oropodes, Oropus, Ouachitychus, Pilactium, Pilopius, Prespelea, Pselaphus, 

Pselaptrichus, Pselaptus, Pseudactium, Pseudotrimium, Pycnoplectus, Pygmactium, Ramecia, 

Reichenbachia, Rhexidius, Rhexius, Rhinoscepsis, Rhybaxis, Saxet, Scalenarthrus, Sebaga, 

Simplona, Sonoma, Speleobama, Stratus, Tetrascapha, Texamaurops, Thesiastes, Thesium, 

Tmesiphorus, Tomplectus, Trichonyx, Trigonoplectus, Trimioarcus, Trimiomelba, Trimioplectus, 

Trisignis, Tychus, Tyrus, Valda, Zonaira. 

Phloeocharinae Erichson 1839: Charhyphus, Dytoscotes, Ecbletus, Phloeocharis, Vicelva. 

Olisthaerinae Thomson 1858: Olisthaerus 

Tachyporinae MacLeay 1825: Bolitobius, Bolitopunctus, Bryophacis, Bryoporus, Carphacis, 

Cilea, Ischnosoma, Lordithon, Mycetoporus, Neobolitobius, Nitidotachinus, Rimulincola, 

Sepedophilus, Tachinomorphus, Tachinus, Tachyporus. 

Trichophyinae Thomson 1858: Trichophya. 

Habrocerinae Mulsant and Rey 1877: Habrocerus. 

Aleocharinae Fleming 1821: Acrimea, Acrotona, Adinopsis, Adota, Agaricochara, 

Agaricomorpha, Aidochara, Alaobia, Aleochara, Aleodorus, Alisalia, Aloconota, Amarochara, 

Amblopusa, Amischa, Anacyptus, Anaduosternum, Anatheta, Anepsiota, Anomognathus, Apalonia, 

Apheloglossa, Apimela, Apteronina, Asthanesita, Atheta, Athetalia, Athetota, Autalia, Bamona, 

Bessobia, Beyeria, Blepharhymenus, Bolitochara, Borboropora, Boreophilia, Boreostiba, 

Brachyusa, Bryobiota, Bryothinusa, Calodera, Canastota, Clusiota, Cordalia, Crataraea, 

Crephalia, Cypha, Cyphea, Datomicra, Decusa, Deinopsis, Devia, Dexiogyia, Diaulota, Diglotta, 

Dimetrota, Dinaraea, Dinardilla, Dinocoryna, Doliponta, Drusilla, Earota, Eburniogaster, 

Ecitocala, Ecitonidia, Ecitoxenida, Elachistarthron, Encephalus, Eumicrota, Euromota, 

Euthorax, Euvira, Falagria, Falagrioma, Falagriota, Fusalia, Gaenima, Geostiba, Gnathusa, 

Gnypeta, Gnypetella, Goniusa, Gymnusa, Gyronycha, Gyrophaena, Halobrecta, Haploglossa, 

Homalota, Homalotusa, Hoplandria, Hydrosmecta, Hydrosmectina, Ilyobates, Iotota, Lamiota, 

Leptagria, Leptobamona, Leptusa, Liogluta, Liparocephalus, Lissagria, Longipeltina, Losiusa,


Lypoglossa, Melanalia, Meotica, Meronera, Micratheta, Micrearota, Microdonia, Microdota, 

Mocyta, Moluciba, Myllaena, Myrmecocephalus, Myrmecopora, Myrmobiota, Myrmoecia, 

Neobeyeria, Nosora, Noverota, Ocalea, Ocyusa, Oligota, Omegalia, Orthodiatelus, Oxypoda, 

Pachycerota, Panalota, Pancota, Paradilacra, Parameotica, Pentanota, Perocalea, Phanerota, 

Phasmota, Philhygra, Philotermes, Phloeopora, Phytosus, Placusa, Platandria, Pontomalota, 

Probeyeria, Pseudomegista, Pseudota, Pesudousipalia, Pulicomorpha, Sableta, Schistacme, 

Schistoglossa, Sibiota, Silusa, Silusida, Sipalia, Sipaliella, Stethusa, Strigota, Strophogastra, 

Symbiochara, Synaptina, Tachyusa, Tarphiota, Teliusa, Termitonidia, Tetradonia, 

Tetralaucopora, Tetrallus, Thamiaraea, Thecturota, Thinusa, Thyasophila, Tinotus, Trachyota, 

Trichiusa, Trichopsenius, Valenusa, Xenistusa, Xenodusa, Xenota, Xesturida, Zyras. 

Trigonurinae Reiche 1865: Trigonurus. 

Scaphidiinae Latreille 1807: Baeocera, Cyparium, Scaphidium, Scaphisoma, Scaphium, 

Toxidium. 

Piestinae Erichson 1839: Hypotelus, Piestus, Siagonium. 

Osoriinae Erichson 1839: Clavilispinus, Eleusis, Lispinus, Nacaeus, Neotrochus, Osorius, 

Renardia, Thoracophorus. 

Oxytelinae Fleming 1821: Aploderus, Apocellus, Bledius, Carpelimus, Coprophilus, Deleaster, 

Manda, Microbledius, Neoxus, Ochthephilus, Oxytelus, Platystethus, Psamathobledius, 

Syntomium, Teropalpus, Thinobius, Thinodromus. 

Oxyporinae Fleming 1821: Oxyporus. 

Megalopsidiinae Leng 1920: Megalopinus. 

Steninae MacLeay 1825: Dianous, Stenus. 

Euaesthetinae Thomson 1859: Cainotyphlus, Edaphus, Euaesthetus, Fenderia, 

Heteroleptophylus, Homeotyphlus, Neoleptotyphlus, Nordenskioeldia, Prototyphlus, Stictocranius, 

Telotyphlus, Xenotyphlus. 

Pseudopsinae Ganglbauer 1895: Asemobius, Nanobius, Pseudopsis, Zalobius. 

Paederinae Fleming 1821: Acalophaena, Achenomorphus, Acrostilicus, Araeocerus, Astenus, 

Biocrypta, Dacnochilus, Echiaster, Homaeotarsus, Lathrobium, Lissobiops, Lithocharis, 

Lobrathium, Medon, Megastilicus, Neomedon, Ochthephilum, Orus, Pachystilicus, Paederus, 

Palaminus, Pinophilus, Rugilus, Scopaeus, Stamnoderus, Stilicolina, Stilicopsis, Stilomedon, 

Sunius, Thinocharis, Xenomedon. 

. Staphylininae Latreille 1802: Acylophorus, Anaquedius, Atanygnathus, Atrecus, Beeria, 

Belonuchus, Bisnius, Cafius, Creophilus, Crinolinus, Diochus, Erichsonius, Gabrius, Gabronthus, 

Gauropterus, Gyrohypnus, Habrolinus, Hadrotes, Hemiquedius, Hesperolinus, Hesperus, 

Heterothops, Hypnogyra, Laetulonthus, Lepitacnus, Leptacinus, Linohesperus, Lissohypnus, 

Lithocharodes, Microlinus, Neobisnius, Neohypnus, Neoxantholinus, Nudobius, Ocypus, 

Ontholestes, Ophiooma, Oxybleptes, Paederopsis, Parothius, Phacophallus, Philonthus, 

Platyprosopus, Quedius, Rabigus, Saurohypnus, Staphylinus, Stenistoderus, Stictolinus, 

Thinopinus, Thyreocephalus, Timagenes, Trigonopselaphus, Tympanophorus, Xantholinus, 

Xanthopygus, Xestolinus, Zenon. 

Not placed in the database: Chionotyphlus, Mitosynum, Salinamexus, Seeversiella. 

Superfamily SCARABAEOIDEA Latreille 1802 

LUCANIDAE Latreille 1804 

Nicaginae Leconte 1861: Nicagus 

Syndesinae MacLeay 1819: Ceruchus, Sinodendron 

Lucaninae Latreille 1804: Dorcus, Lucanus, Platyceroides, Platyceropsis, Platycerus, 

Pseudolucanus 

PASSALIDAE Leach 1815 

Passalinae Leach 1815: Passalus, Popilius. 

TROGIDAE MacLeay 1819 

Trox. 

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CLASSIFICATION

26 



GLARESIDAE Semenov 1932 

Glaresis. 

PLEOCOMIDAE Leconte 1861 

Acoma, Benedictia, Pleocoma. 

DIPHYLLOSTOMATIDAE Holloway 1972 

Diphyllostoma. 

GEOTRUPIDAE Latreille 1802 

Bolboceratinae Mulsant 1842: Bolbelasmus, Bolboceras, Bolbocerastes, Bolbocerosoma, 

Bolborhombus, Bradycinetulus, Eucanthus. 

Geotrupinae Latreille 1802: Ceratophyus, Geotrupes, Mycotrupes, Peltotrupes. 

OCHODAEIDAE Mulsant and Rey 1871 

Ochodaeus, Pseudochodaeus. 

CERATOCANTHIDAE Cartwright and Gordon 1971 

Ceratocanthus, Germarostes. 

HYBOSORIDAE Erichson 1847 

Pachyplectrus, Phaeochrous. 

GLAPHYRIDAE MacLeay 1819 

Lichnanthe. 

SCARABAEIDAE Latreille 1802 

Aphodiinae Leach 1815: Aegialia, Annegialia, Aphodius, Aphotaenius, Ataenius, Caelius, 

Dialytellus, Dialytes, Diastictus, Euparixia, Euparia, Geopsammodius, Leiopsammodius, 

Micraegilia, Neopsammodius, Odontopsammodius, Oxyomus, Platytomus, Pleurophorus, 

Psammodius, Pseudataenius, Rhyssemus, Tesarius, Trichiorhyssemus, Xeropsamobeus. 

Scarabaeinae Latreille 1802: Ateuchus, Boreocanthon, Canthon, Copris, Deltochilum, 

Megathopa, Oniticellus, Onthophagus, Phanaeus, Pinotus. 

Melolonthinae MacLeay 1819: Amphimallon, Ceraspis, Chaunocolus, Chnaunanthus, 

Coenonycha, Dichelonyx, Dinacoma, Diplotaxis, Fossocarus, Gronocarus, Gymnopyge, Hoplia, 

Thyce, Hypotrichia, Isonychus, Leptohoplia, Macrodactylus, Maladera, Nefoncerus, Oncerus, 

Parathyce, Phobetus, Phyllophaga, Plectrodes, Podolasia, Polylamina, Polyphylla, Serica, Thyce. 

Rutelinae MacLeay 1819: Anomala, Anomalacra, Chrysina, Cotalpa, Parabyrsopolis, 

Paracotalpa, Parastasia, Pelidnota, Plusiotis, Popillia, Pseudocotalpa, Rutela, Strigoderma. 

Dynastinae MacLeay 1819: Ancognatha, Aphonides, Aphonus, Archophileurus, Aspidolea, 

Coscinocephalus, Cyclocephala, Dynastes, Dyscinetus, Euetheola, Hemiphileurus, Ligyrus, 

Megasoma, Orizabus, Phileurus, Stenocrates, Strategus, Xyloryctes. 

Cetoniinae Leach 1815: Anatropis, Chlorixantha, Cineretis, Cotinis, Cremastocheilus, 

Epicometis, Euphoria, Genuchinus, Gnorimella, Gymnetis, Lissomelas, Osmoderma, Potosia, 

Psilocnemis, Stephanucha, Trichiotinus, Trigonopeltastes, Valgus. 

Superfamily SCIRTOIDEA Fleming 1821 

EUCINETIDAE Lacordaire 1857 

Eucinetus, Euscaphurus. 

CLAMBIDAE Fisher 1821 

Calyptomerinae Crowson 1955: Calyptomerus. 

Clambinae Fisher 1821: Clambus, Loricaster. 

SCIRTIDAE Fleming 1821 

Cyphon, Elodes, Microcara, Ora, Prionocyphon, Sarabandus, Scirtes. 

Superfamily DASCILLOIDEA Guerin-Meneville 1843


DASCILLIDAE Guerin-Meneville 1843 

Dascillinae Guerin-Meneville 1843: Dascillus. 

Karumiinae Escalera 1913: Anorus. 

RHIPICERIDAE Latreille 1834 

Sandalus. 

Superfamily BUPRESTOIDEA Leach 1815 

BUPRESTIDAE Leach 1815 

Schizopodinae Leconte 1861: Dystaxia, Glyptoscelimorpha, Schizopus. 

Buprestinae Leach 1815: Acmaeodera, Acmaeoderoides, Acmaeoderopsis, Actenodes, 

Agaeocera, Anambodera, Ancylotela, Anthaxia, Beerellus, Buprestis, Chalcophora, 

Chrysobothris, Chrysophana, Descarpentriesina, Dicerca, Haplostethus, Hesperorhipis, 

Hippomelas, Juniperella, Melanophila, Nanularia, Oxypteris, Polycesta, Psiloptera, Ptosima, 

Spectralia, Texania, Thrincopyge, Trachykele, Trichinorhipis, Trigonogya, Tyndaris, 

Xenomelanophila, Xenorhipis. 

Argilinae Laporte 1835: Agrilus, Brachys, Eupristocerus, Pachyschelus, Paragrilus, 

Taphrocerus. 

Superfamily BYRRHOIDEA Latreille 1804 

BYRRHIDAE Latreille 1804 

Byrrhinae Latreille 1804: Byrrhus, Cytillus, Eusomalia, Exomella, Lioligus, Lioon, Listemus, 

Morychus, Porcinolus, Simplocaria, Tylicus. 

Syncalyptinae Mulsant and Rey 1869: Curimopsis, Sierraclava, Syncalypta. 

Amphicyrtinae Leconte 1861: Amphicyrta. 

ELMIDAE Curtis 1830 

Larainae Leconte 1861: Lara, Phanocerus. 

Elminae Curtis 1830: Ampumixis, Ancyronyx, Atractelmis, Cleptelmis, Cylloepus, Dubiraphia, 

Gonielmis, Heterelmis, Heterlimnius, Hexacylloepus, Huleechius, Macrelmis, Macronychus, 

Microcylloepus, Narpus, Neocylloepus, Neoelmis, Optiservus, Ordobrevia, Oulimnius, 

Promoresia, Rhizelmis, Stenelmis, Xenelmis, Zaitzevia. 

DRYOPIDAE Billberg 1820 

Dryops, Helichus, Pelonomus, Posthelichus, Stygoparnus. 

LUTROCHIDAE Kasap and Crowson 1975 

Lutrochus. 

LIMNICHIDAE Erichson 1846 

Limnichinae Erichson 1846: Eulimnichus, Lichminus, Limnichites, Limnichoderus, 

Physemus. 

Cephalobyrrhinae Champion 1925: Throscinus. 

HETEROCERIDAE MacLeay 1825 

Heterocerinae MacLeay 1825: Augyles, Centuriatus, Efflagitatus, Explorator, Heterocerus, 

Lanternarius, Lapsus, Microaugyles, Neoheterocerus, Peditatus, Tropicus. 

PSEPHENIDAE Lacordaire 1854 

Eubriinae Lacordaire 1857: Acneus, Alabameubria, Dicranopselaphus, Ectopria. 

Eubrianacinae Jacobson 1913: Eubrianax. 

Psepheninae Lacordaire 1854: Psephenus. 

PTILODACTYLIDAE Laporte 1836 

Anchytarsinae Champion 1897: Anchycteis, Anchytarsus. 

Cladotominae Pic 1914: Odontonyx. 

Araeopidiinae Lawrence 1991: Araeopidius. 

CHELONARIIDAE Blanchard 1845 

Chelonarium. 

27 

CLASSIFICATION

28 

EULICHADIDAE Crowson 1973 

Stenocolus. 

CALLIRHIPIDAE Emden 1924 

Zenoa. 



Superfamily ELATEROIDEA Leach 1815 

ARTEMATOPODIDAE Lacordaire 1857 

Artematopodinae Lacordaire 1857: Allopogonia, Eurypogon, Macropogon. 

BRACHYPSECTRIDAE Leconte and Horn 1883 

Brachypsectra. 

CEROPHYTIDAE Latreille 1834 

Cerophytum. 

EUCNEMIDAE Eschscholtz 1829 

Perothopinae Lacordaire 1857: Perothops. 

Phyllocerinae Reitter 1905: Anelastes. 

Pseudomeninae Muona 1993: Schizophyllus. 

Palaeoxeninae Muona 1993: Palaeoxenus. 

Melasinae Fleming 1821: Adelothyreus, Arrhipis, Diphytaxis, Entomophthalmus, Epiphanis, 

Hylis, Hylochares, Isorhipis, Melasis, Microrhagus, Nematodes, Rhagomicrus, Sarpedon, 

Xylophilus. 

Eucneminae Eschscholtz 1829: Deltometopus, Dendrocharis, Eucnemis, Poecilochrus, 

Proutianus, Stethon, Vitellius. 

Macraulacinae Fleutiaux 1923: Euryptychus, Fornax. 

THROSCIDAE Laporte 1840 

Aulonothroscus, Pactopus, Trixagus. 

ELATERIDAE Leach 1815 

Cebrioninae Latreille 1802: Anthracopteryx, Aplastus, Euthysanius, Octinodes, Scaptolenus, 

Selonodon. 

Lissominae Laporte 1835: Drapetes, Oestodes. 

Semiotinae Jacobson 1913: Oistus. 

Pityobiinae Hyslop 1917: Pityobius. 

Agrypninae Candeze 1857: Aeolus, Agrypnus, Alaus, Chalcolepidius, Conoderus, Danosoma, 

Hemirhipus, Heteroderes, Lacon, Lanelater, Meristhus, Pyrophorus, Rismethus. 

Denticollinae Stein and Weise 1877: Athous, Berninelsonius, Bladus, Ctenicera, Denticollis, 

Eanus, Elathous, Fleutiauxellus, Harminius, Hemicrepidius, Hypnoidus, Hypolithus, 

Leptoschema, Ligmargus, Limonius, Margaiostus, Melanactes, Micrathous, Migiwa, 

Neohypdonus, Paradonus, Paroedostethus, Perissarthron, Proludius, Pseudomelanactes. 

Negastriinae Nakane and Kishii 1956: Negastrius, Oedostethus, Zorochrus. 

Elaterinae Leach 1815: Agriotella, Agriotes, Ampedus, Anchastus, Atractodes, Blauta, Dalopius, 

Dicrepidius, Diplostethus, Dipropus, Elater, Glyphonyx, Ischiodontus, Megapenthes, Melanotus, 

Neotrichophorus, Orthostethus, Oxygonus, Parallelostethus, Physorhinus, Sericus. 

Cardiophorinae Candeze 1859: Aphricus, Aptopus, Cardiophorus, Coptostethus, Esthesopus, 

Horistonotus. 

Unplaced in the database: Ascoliocerus. 

LYCIDAE Laporte 1836 

Lycinae Laporte 1836: Caenia, Calopteron, Celetes, Idiopteron, Lycus. 

Erotinae Leconte 1881: Dictyopterus, Eropterus, Eros, Lopheros, Plateros, Platycis, 

Pseudoplateros. 

Calochrominae Lacordaire 1857: Adoceta, Caloptognatha, Lucaina, Lygistopterus. 

TELEGEUSIDAE Leng 1920 

Telegeusis.


PHENGODIDAE Leconte 1861 

Phengodinae Leconte 1861: Cenophengus, Euryopa, Mastinocerus, Paraptorthodius, 

Phengodes, Zarhipis. 

LAMPYRIDAE Latreille 1817 

Pterotinae Leconte 1861: Pterotus. 

Cyphonocerinae Crowson 1972: Pollaclasis. 

Ototretinae McDermott 1964: Brachylampis. 

Lampyrinae Latreille 1817: Aspisoma, Ellychnia, Lampyris, Lucidota, Micronaspis, 

Microphotus, Paraphausis, Photinus, Pleotomus, Pyractonema, Pyropyga, Tenaspis. 

Photurinae Lacordaire 1857: Bicellonycha, Phausis, Photuris. 

OMETHIDAE Leconte 1861 

Matheteinae Leconte 1881: Ginglymocladus, Matheteus. 

Omethinae Leconte 1861: Blatchleya, Malomethes, Omethes, Symphyomethes, Troglomethes. 

CANTHARIDAE Imhoff 1856 

Cantharinae Imhoff 1856: Absidiella, Ancistronycha, Cantharis, Cultellunguis, Cyrtomoptera, 

Hemipodistra, Podabrus, Pseudoabsidia, Rhagonycha, Raxonycha. 

Silinae Mulsant 1862: Discodon, Plectonotum, Polemius, Silis, Tytthonyx. 

Malthininae Kiesenwetter 1852: Caccodes, Frostia, Malthinus, Malthodes. 

Chauliognathinae Leconte 1861: Belotus, Chauliognathus, Ichthyurus, Trypherus. 

Unplaced in the database: Hatchiana. 

Superfamily DERODONTOIDEA Leconte 1861 

DERODONTIDAE Leconte 1861 

Peltasticinae Leconte 1861: Peltastica. 

Derodontinae Leconte 1861: Derodontus. 

Laricobiinae Mulsant and Rey 1864: Laricobius. 

Superfamily BOSTRICHOIDEA Latreille 1802 

NOSODENDRIDAE Erichson 1846 

Nosodendron. 

DERMESTIDAE Latreille 1804 

Dermestinae Latreille 1804: Dermestes. 

Thorictinae Agassiz 1846: Thorictodes. 

Orphilinae Leconte 1861: Orphilus. 

Trinodinae Casey 1900: Apsectus. 

Thylodriinae Semenov 1913: Thylodrias. 

Attageninae Laporte 1840: Attagenus, Novelsis. 

Megatominae Leach 1815: Anthrenus, Cryptorhopalum, Ctesias, Dearthrus, Megatoma, 

Orphinus, Pseudohadrotoma, Trogoderma. 

ENDECATOMIDAE Leconte 1861 

Endecatomus. 

BOSTRICHIDAE Latreille 1802 

Polycaoninae Lesne 1896: Melalgus, Polycaon. 

Bostrichinae Latreille 1802: Amphicerus, Apatides, Bostrichus, Dendrobiella, Dinapate, 

Lichenophanes, Micrapate, Scobicia, Sinoxylon, Tetrapriocera, Xylobiops, Xyloblaptus, 

Xylomeira. 

Psoinae Blanchard 1851: Psoa. 

Dinoderinae Thomson 1863: Dinoderus, Prostephanus, Rhyzopertha, Stephanopachys. 

Lyctinae Billberg 1820: Lyctoxylon, Lyctus, Minthea, Togoxylon. 

29 

CLASSIFICATION

30 



ANOBIIDAE Fleming 1821 

Eucradinae Leconte 1861: Eucrada, Hedobia, Neohedobia, Ptilineurus. 

Ptininae Latreille 1802: Eupauloecus, Gibbium, Gnostus, Mezium, Niptinus, Niptus, Pitnus, 

Ptinus, Sphaericus, Trigonogenius. 

Ernobiinae Pic 1912: Episernus, Ernobius, Microzogus, Ozognathus, Paralobium, Utobium, 

Xarifa, Xestobium. 

Anobiinae Fleming 1821: Actenobius, Anobiopsis, Anobium, Colposternus, Ctenobium, 

Desmatogaster, Euceratocerus, Falsogastrallus, Gastrallus, Hadrobregmus, Hemicoelus, 

Microbregma, Nicobium, Oligomerus, Platybregmus, Priobium, Stegobium, Trichodesma, 

Xeranobium. 

Ptilininae Shuckard 1840: Ptilinus. 

Xyletininae Gistel 1856: Euvrilletta, Lasioderma, Megorama, Vrilletta, Xyletinus, Xyletomerus. 

Dorcatominae Thomson 1859: Byrrhodes, Caenocara, Calymmaderus, Cryptoramorphus, 

Dorcatoma, Petalium, Protheca, Sculptotheca, Stagetus, Stichtoptychus, Striatheca. 

Mesocoelopodinae Mulsant and Rey 1864: Cryptorama, Mesocoelopus, Tricorynus. 

Superfamily LYMEXYLOIDEA Fleming 1821 

LYMEXYLIDAE Fleming 1821 

Hylecoetinae Gistel 1856: Hylecoetus. 

Lymexylinae Fleming 1821: Atractocerus. 

Melittommatinae Wheeler 1986: Melittomma. 

Superfamily CLEROIDEA Latreille 1802 

TROGOSSITIDAE Latreille 1802 

Peltinae Kirby 1837: Ostoma, Thymalus. 

Lophocaterinae Crowson 1964: Eronyxa, Grynocharis, Lophocateres, Lycoptis. 

Calitinae Reitter 1922: Calitys. 

Trogossitinae Latreille 1802: Airora, Corticotomus, Cylidrella, Euschaefferia, Nemozoma, 

Temnocheila, Tenebroides. 

CLERIDAE Latreille 1802 

Thaneroclerinae Chapin 1924: Ababa, Thaneroclerus, Zenodosus. 

Tillinae Leach 1815: Araeodontia, Bostrichoclerus, Callotillus, Cymatodera, Lecontella, 

Monophylla, Perilypus. 

Hydnocerinae Spinola 1844: Isohydnocera, Phyllobaenus, Wolcottia. 

Clerinae Latreille 1802: Aulicus, Colyphus, Enoclerus, Opilo, Placopterus, Thanasimus, 

Trichodes. 

Epiphloeinae Kuwert 1893: Ichnea, Neichnea, Phlogistosternus. 

Enopliinae Gistel 1856: Chariessa, Cregya, Neorthopleura, Pelonides, Pelonium. 

Tarsosteninae Jacquelin du Val 1860: Tarsostenus. 

Korynetinae Laporte 1836: Lebasiella, Loedelia, Necrobia, Opetiopalpus. 

MELYRIDAE Leach 1815 

Melyrinae Leach 1815: Eurelymis, Melyrodes. 

Rhadalinae Leconte 1861: Dasyrhadus, Rhadalus, Semijulistus. 

Dasytinae Laporte 1840: Adasytes, Amecocerus, Amphivectura, Asydates, Byturosoma, 

Cradytes, Dasytastes, Dasytellus, Dasytes, Dolichosoma, Emmenotarsis, Enallonyx, 

Eschatocrepis, Eudasytes, Eutricholistra, Eutrichopleurus, Hoppingiana, Leptovectura, 

Listrimorpha, Listromimus, Listropsis, Mecomycter, Neodasytes, Paradasytes, Pristoscelis, 

Pseudallonyx, Pseudasydates, Sydates, Trichochronellus, Trichochrous, Vectura, Vecturoides.


Malachiinae Fleming 1821: Ablechrus, Anthocomus, Attalus, Attalusinus, Chaetocoelus, Collops, 

Ebaeus, Endeodes, Hypebaeus, Malachius, Microlipus, Nodopus, Tanaops, Temnopsophus, 

Trophimus. 

Superfamily CUCUJOIDEA Latreille 1802 

SPHINDIDAE Jacquelin du Val 1860 

Odontosphindinae Sen Gupta and Crowson 1979: Odontosphindus. 

Sphindinae Jacquelin du Val 1860: Eurysphindus, Sphindus. 

BRACHYPTERIDAE Erichson 1845 

Amartus, Anthonaeus, Boreades, Brachypterolus, Brachypterus, Heterhelus, Kateretes, 

Neobrachypterus. 

NITIDULIDAE Latreille 1802 

Carpophilinae Erichson 1842: Amphicrossus, Carpophilus, Epuraea. 

Meligethinae Thomson 1859: Meligethes. 

Nitidulinae Latreille 1802: Amphotis, Aphenolia, Camptodes, Cychramus, Cyllodes, Haptoncus, 

Lobiopa, Nitidula, Omosita, Orthopeplus, Oxycnemus, Pallodes, Phenolia, Pocadius, Prometopia, 

Pseudothalycra, Psilopyga, Soronia, Stelidota, Thalycra. 

Cillaeinae Kirejtshuk and Audisio 1986: Brachypeplus, Colopterus, Conotelus. 

Cryptarchinae Thomson 1859: Cryptarcha, Glischrochilus, Pityophagus. 

Cybocephalinae Jacquelin du Val 1858: Cybocephalus. 

SMICRIPIDAE Horn 1879 

Smicrips. 

MONOTOMIDAE Laporte 1840 

Rhizophaginae Redtenbacher 1845: Rhizophagus. 

Monotominae Laporte 1840: Bactridium, Europs, Hesperobaenus, Macreurops, Monotoma, 

Leptipsius, Phyconomus, Pycnotomina. 

SILVANIDAE Kirby 1837 

Brontinae Erichson 1845: Psammoecus, Telephanus, Uleiota. 

Silvaninae Kirby 1837: Ahasverus, Cathartosilvanus, Cathartus, Monanus, Nausibius, 

Oryzaephilus, Pensus, Planismus, Silvanus. 

PASSANDRIDAE Erichson 1845 

Catogenus, Prostomis, Taphroscelidia. 

CUCUJIDAE Latreille 1802 

Cucujinae Latreille 1802: Cucujus, Pediacus. 

LAEMOPHLOEIDAE Ganglbauer 1899 

Charaphloeus, Cryptolestes, Deinophloeus, Dysmerus, Laemophloeus, Lathropus, Metaxyphloeus, 

Narthecius, Placonotus, Rhinomalus. 

PHALACRIDAE Leach 1815 

Phalacrinae Leach 1815: Acylomus, Gorginus, Leptostilbus, Litochropus, Litochrus, Litolibrus, 

Ochrolitus, Olibroporus, Olibrus, Phalacropsis, Phalacrus, Stilbus, Tinodemus. 

CRYPTOPHAGIDAE Kirby 1837 

Hypocoprinae Reitter 1879: Hypocoprus. 

Cryptophaginae Kirby 1837: Antherophagus, Caenoscelis, Cryptophagus, Henoticus, 

Henotiderus, Myrmedophila, Pteryngium, Salebius, Telmatophilus. 

Atomariinae Leconte 1861: Anichera, Atomaria, Ephistemus, Parephistemus, Tisactia. 

LANGURIIDAE Crotch 1873 

Xenoscelinae Ganglbauer 1899: Hapalips, Loberus, Pharaxontha. 

Languriinae Crotch 1873: Acropteroxys, Dasydactylus, Languria, Langurites. 

Cryptophilinae Casey 1900: Cryptophilus. 

31 

CLASSIFICATION

32 



Toraminae Sen Gupta 1967: Toramus. 

EROTYLIDAE Latreille 1802 

Dacninae Gistel 1856: Dacne. 

Megalodacninae Sen Gupta 1969: Megalodacne, Microsternus. 

Tritominae Curtis 1834: Haematochiton, Hirsutotriplax, Ischyrus, Mycotretus, Pseudischyrus, 

Triplax, Tritoma. 

Erotylinae Latreille 1802: Cypherotylus. 

BYTURIDAE Jacquelin du Val 1858 

Byturinae Jacquelin du Val 1858: Byturus, Xerasia. 

BIPHYLLIDAE Leconte 1861 

Anchorius, Diplocoelus. 

BOTHRIDERIDAE Erichson 1845 

Teredinae Seidlitz 1888: Oxylaemus. 

Anommatinae Ganglbauer 1899: Anommatus. 

Bothriderinae Erichson 1845: Bothrideres, Deretaphrus, Erotylathris, Lithophorus, Sosylus. 

CERYLONIDAE Billberg 1820 

Euxestinae Grouvelle 1908: Hypodacne. 

Ostomopsinae Sen Gupta and Crowson: Ostomopsis. 

Murmidiinae Jacquelin du Val 1858: Botrodus, Murmidius. 

Ceryloninae Billber 1820: Cerylcautomus, Cerylon, Lapethus, Mychocerus, Philothermus. 

ENDOMYCHIDAE Leach 1815 

Holoparamecinae Seidlitz 1888: Holoparamecus. 

Eupsilobiinae Casey 1895: Eidoreus. 

Endomychinae Leach 1815: Endomychus. 

Epipocinae Gorham 1873: Danae, Epipocus, Stenotarsus. 

Lycoperdininae Redtenbacher 1844: Aphorista, Lycoperdina, Mycetina, Phymaphora, Rhanidea. 

Mycetaeinae Jacquelin du Val 1857: Mycetaea. 

Anamorphinae Strohecker 1953: Anamorphus, Bystus, Clemmus, Rhymbomicrus, Stethorhanis, 

Symbiotes. 

Xenomycetinae Strohecker 1962: Xenomycetes. 

COCCINELLIDAE Latreille 1807 

Sticholotidinae Weise 1901: Cephaloscymnus, Coccidophilus, Delphastus, Gnathoweisea, 

Microweisea, Nipus. 

Coccidulinae Mulsant 1846: Anovia, Azya, Coccidula, Exoplectra, Pseudoazaya, Rhodolia, 

Rhyzobius. 

Scymninae Mulsant 1846: Blaisdelliana, Brachiacantha, Cryptognatha, Cryptolaemus, Didion, 

Diomus, Helesius, Hyperaspidius, Hyperaspis, Nephaspis, Nephus, Scymnus, Selvadius, Stethorus, 

Thalassa, Zagloba, Zilus. 

Chilocorinae Mulsant 1846: Arawana, Axion, Brumoides, Chilocorus, Exochomus, Halmus. 

Coccinellinae Latreille 1807: Aphidecta, Anatis, Anisosticta, Adalia, Calvia, Ceratomegilla, 

Coccinella, Coelophora, Coleomegilla, Cycloneda, Harmonia, Hippodamia, Macroanaemia, 

Mulsantina, Myzia, Naemia, Neoharmonia, Olla, Paranaemia, Propylaea, Psyllobora. 

Epilachninae Mulsant 1846: Epilachna, Subcoccinella. 

CORYLOPHIDAE Leconte 1852 

Corylophinae Leconte 1852: Aenigmaticum, Bathona, Corylophodes, Eutrilia, Foadia, 

Gronevus, Orthoperus, Ryobius. 

Sericoderinae Matthews 1888: Sericoderus. 

Parmulinae Poey 1854: Arthrolips, Molamba, Sacium. 

LATRIDIIDAE Erichson 1842 

Latridiinae Erichson 1842: Adistemia, Akalyptoischion, Aridius, Cartodere, Enicmus, Eufallia, 

Latridius, Metophthalmus, Microgramme, Revelieria, Stephostethus, Thes. 

Corticariinae Curtis 1829: Corticaria, Corticarina, Fuchsina, Melanophthalma.


Superfamily TENEBRIONOIDEA Latreille 1802 

MYCETOPHAGIDAE Leach 1815 

Mycetophaginae Leach 1815: Lendomus, Litargus, Mycetophagus, Myrmechixenis, Thrimolus, 

Typhaea. 

Bergininae Leng 1920: Berginus. 

ARCHEOCRYPTICIDAE Kaszab 1964 

Enneboeus. 

CIIDAE Leach 1819 

Sphindociinae Lawrence 1974: Sphindocis. 

Ciinae Leach 1819: Ceracis, Cis, Dolichocis, Ennearthron, Hadraule, Malacocis, Octotemnus, 

Orthocis, Plesiocis, Rhopalodontus, Strigocis, Sulcacis. 

TETRATOMIDAE Billberg 1820 

Piseninae Miyatake 1960: Pisenus, Triphyllia. 

Tetratominae Billberg 1820: Abstrulia, Incolia, Tetratoma. 

Penthinae Lacordaire 1859: Penthe. 

MELANDRYIDAE Leach 1815 

Hallomeninae Mulsant 1856: Hallomenus. 

Eustrophinae Gistel 1856: Eustrophinus, Eustrophus, Holostrophus, Synstrophus. 

Melandryinae Leach 1815: Abdera, Amblyctis, Anisoxya, Dircaea, Enchodes, Hypulus, Lederia, 

Melandrya, Microscapha, Microtonus, Orchesia, Phloeotrya, Phryganophilus, Polypria, 

Prothalpia, Rushia, Scotochroa, Scotochroides, Serropalpus, Spilotus, Xylita, Zilora. 

Osphyinae Mulsant 1856: Osphya. 

MORDELLIDAE Latreille 1802 

Mordellinae Latreille 1802: Conalia, Glipa, Glipodes, Glipostenoda, Hoshihananomia, 

Isotrilophus, Mordella, Mordelistena, Paramordellaria, Tolidomordella, Tomoxia, 

Yakuhananomia. 

RHIPIPHORIDAE Gemminger and Harold 1870 

Pelecotominae Seidlitz 1875: Pelecotoma, Trigonodera. 

Ptilophorinae Gerstaecker 1855: Toposcopus. 

Rhipidiinae Gerstaecker 1855: Ripidius. 

Rhipiphorinae Gemminger and Harold 1870: Macrosiagon, Rhipiphorus. 

COLYDIIDAE Erichson 1842 

Pycnomerinae Erichson 1845: Pycnomerus. 

Colydiinae Erichson 1842: Acolobicus, Anchomma, Aulonium, Bitoma, Colydium, Coxelus, 

Denophloeus, Endeitoma, Eucicones, Eudesmula, Eulachus, Lasconotus, Lobogestoria, 

Megataphrus, Microsicus, Monoedus, Myrmechixenus, Namunaria, Nematidium, Phloeonemus, 

Pseudotaphrus, Rhagodera, Rustleria, Synchita. 

MONOMMATIDAE Blanchard 1845 

Aspathenes, Hyporhagus. 

ZOPHERIDAE Solier 1834 

Zopherinae Solier 1834: Noserus, Phellopsis, Phloeodes, Zopherus. 

Usechinae Horn 1867: Usechimorpha, Usechus. 

TENEBRIONIDAE Latreille 1802 

Lagriinae Latreille 1825: Arthromacra, Statira. 

Phrenapatinae Solier 1834: Clamoris, Dioedus 

Pimeliinae Latreille 1802: Anepsius, Araeoschizus, Armalia, Asida, Asidina, Asidopsis, Astrotus, 

Auchmobius, Batuliodes, Batuliomorpha, Batulius, Bothrotes, Centrioptera, Chilometopon, 

Cnemodinus, Coelus, Conoecus, Craniotus, Cryptadius, Cryptoglossa, Cyrtomius, Dacoderus, 

Edrotes, Emmenastus, Epitragodes, Eschatomoxys, Eschatoporis, Eurymetopon, Glyptasida, 

33 

CLASSIFICATION

34 



Gonasida, Hylocrinus, Lobometopon, Megasida, Melanastus, Metopoloba, Metoponium, 

Micromes, Microschatia, Nyctoporis, Oxygonodera, Parasida, Pechalius, Phegoneus, Philolithus, 

Polemiotus, Schizillus, Schoenicus, Stenomorpha, Stenosides, Stethasida, Steriphanus, Stibia, 

Threnus, Telabis, Trimytis, Trichiasida, Trichiotes, Triorophus, Typhlusechus. 

Tenebrioninae Latreille 1802: Aconobius, Adelina, Adelonia, Alaephus, Alaudes, Alphitobius, 

Ammodonus, Anaedus, Apocrypha, Argoporis, Biomorphus, Bius, Blaps, Blapstinus, 

Bolitophagus, Bolitopherus, Branchus, Centronopus, Cerenopus, Cheirodes, Cnemeplatia, 

Conibiosoma, Conibius, Coniontis, Conisattus, Cybotus, Cyclosattus, Cymatothes, Cynaeus, 

Doliodesmus, Eleates, Eleodes, Eleodimorpha, Embaphion, Ephalus, Eulabis, Eupsophulus, 

Eusattus, Eutochia, Gonocephalum, Helops, Hypogena, Idiobates, Lariversius, Latheticus, 

Leichenum, Lyphia, Mecysmus, Megeleates, Meracantha, Mycotrogus, Nautes, Neatus, 

Neobaphion, Nocibiotes, Notibius, Opatrinus, Palorus, Paratenetus, Prateus, Pseudephalus, 

Rhipidandrus, Scotobaenus, Sitophagus, Stenotrichus, Tarpela, Tenebrio, Tharsus, Tonibius, 

Tribolium, Trichoton, Trogloderus, Uloma, Ulus, Zophobas. 

Alleculinae Laporte 1840: Aletha, Anamphidora, Andrimus, Androchirus, Capnochroa, 

Charisius, Hymenochara, Hymenorus, Isomira, Knausia, Lobopoda, Lystronichus, Menoeceus, 

Mycetochara, Onychomira, Phedius, Prostenus, Pseudocistela, Stenochidus, Tedinus, Telesicles. 

Diaperinae Latreille 1802: Alphitophagus, Corticeus, Crypticus, Diaperis, Gnathocerus, 

Liodema, Metaclisa, Neomida, Palembus, Pentaphyllus, Phaleria, Platydema, Scaphidema, 

Trachyscelis, Ulomoides. 

Coelometopinae Lacordaire 1859: Alobates, Apsida, Cibdelis, Coelocnemis, Glyptotus, 

Haplandrus, Iphthiminus, Merinus, Oenopion, Polopinus, Polypleurus, Strongylium, Talanus, 

Upis, Xylopinus. 

Not placed in the database: Alegoria, Calymmus, Melasmana. 

PROSTOMIDAE Thomson 1859 

Prostomis. 

SYNCHROIDAE Lacordaire 1859 

Synchroa, Mallodrya 

OEDEMERIDAE Latreille 1810 

Calopodinae Costa 1852: Calopus, Sparedrus. 

Oedemerinae Latreille 1810: Asclera, Copidita, Ditylus, Eumecomera, Heliocis, Hypasclera, 

Nacerdes, Oxacis, Oxycopis, Paroxacis, Rhinoplatia, Sisenes, Vasaces, Xanthochroa, 

Xanthochroina. 

STENOTRACHELIDAE Thomson 1859 

Stenotrachelinae Thomson 1859: Anelpistus, Stenotrachelus. 

Nematoplinae Leconte 1862: Nematoplus. 

Cephaloinae Leconte 1862: Cephaloon. 

MELOIDAE Gyllenhal 1810 

Meloinae Gyllenhal 1810: Cordylospasta, Cysteodemus, Epicauta, Eupompha, Linsleya, Lytta, 

Megetra, Meloe, Phodaga, Pleuropasta, Pyrota, Tegrodera, Thambospasta, 

Nemognathinae Laporte 1840: Gnathium, Hornia, Nemognatha, Pseudozonitis, 

Rhyphonemognatha, Tetraonyx, Tricrania, Zonitis. 

MYCTERIDAE Blanchard 1845 

Mycterinae Blanchard 1845: Mycterus. 

Lacconotinae Leconte 1862: Lacconotus. 

Hemipeplinae Lacordaire 1854: Hemipeplus. 

BORIDAE Thomson 1859 

Borinae Thomson 1859: Boros, Lecontia. 

PYTHIDAE Solier 1834 

Priognathus, Pytho, Sphalma, Trimitomerus. 

PYROCHOIDAE Latreille 1807 

Tydessinae Nikitsky 1986: Tydessa.


Pedilinae Lacordaire 1859: Anisotria, Neopedilus, Pedilus. 

Pyrochorinae Latreille 1807: Dendroides, Neopyrochroa, Schizotus. 

Agnathinae Lacordaire 1859: Cononotus. 

SALPINGIDAE Leach 1815 

Othniinae Leconte 1861: Elacatis. 

Agleninae Horn 1878: Aglenus. 

Inopeplinae Grouvelle 1908: Inopeplus. 

Salpinginae Leach 1815: Rhinosimus, Sphaeriestes, Vincenzellus. 

Aegialitinae Leconte 1862: Aegialites. 

Dacoderinae Leconte 1862: Dacoderus. 

ANTHICIDAE Latreille 1819 

Eurygeniinae Leconte 1862: Bactrocerus, Duboisius, Eurygenius, Mastoremus, Neoeurygenius, 

Quadrius, Rilettius, Stereopalpus. 

Macratriinae Leconte 1862: Macratria. 

Ischaliinae Blair 1920: Ischalia. 

Tomoderinae Bonadona 1961: Tomoderus. 

Anthicinae Latreille 1819: Acanthinus, Amblyderus, Anthicus, Euvacusus, Formicilla, 

Ischyropalpus, Leptanthicus, Leptoremus, Malporus, Mecynotarsus, Notoxus, Sapintus, 

Tanarthrus, Thicanus, Vacusus. 

ADERIDAE Winkler 1927 

Aderus, Ariotus, Axylophilus, Cnopus, Emelinus, Ganascus, Gymnoganascus, Pseudariotus, 

Vanonus. 

SCRAPTIIDAE Mulsant 1856 

Scraptiinae Mulsant 1856: Canifa, Neoscraptia, Pectotoma, Scraptia. 

Anaspidinae Mulsant 1856: Allopoda, Anaspis, Diclidia, Larisia, Nassipa, Naucles, Pentaria, 

Silaria, Sphingocephalus. 

Superfamily CHRYSOMELOIDEA Latreille 1802 

CERAMBYCIDAE Latreille 1802 

Disteniinae Thomson 1860: Distenia. 

Paradrinae Blanchard 1845: Parandra. 

Prioninae Latreille 1802: Archodontes, Derancistrus, Derobrachus, Ergates, Neomallodon, 

Orthosoma, Prionus, Stenodontes, Strongylaspis, Tragosoma. 

Spondylidinae Serville 1832: Arhopalus, Asemum, Atimia, Megasemum, Michthisoma, 

Paratimia, Scaphinus, Spondylis, Tetropium. 

Necydalinae Latreille 1825: Necydalis, Ulochaetes. 

Lepturinae Latreille 1802: Acmaeops, Acmaeopsoides, Alosternida, Analeptura, Anastrangalia, 

Anthophylax, Bellamira, Brachyleptura, Brachysomida, Centrodera, Charisalia, Choriolaus, 

Comacmaeops, Cortodera, Cosmosalia, Cyphonotida, Desmocerus, Dorcasina, Encyclops, 

Evodinus, Gaurotes, Gnathacmaeops, Grammoptera, Idiopidonia, Judolia, Leptalia, 

Leptorhabdium, Leptura, Lepturopsis, Lycochoriolaus, Megachoriolaus, Metacmaeops, 

Neanthophylax, Neoalosterna, Neobellamira, Orthochoriolaus, Ortholeptura, Pachyta, Pidonia, 

Piodes, Pseudogaurotina, Pseudostranglia, Pygoleptura, Pyrotrichus, Rhagium, Sachalinobia, 

Stenocorus, Stenostrophia, Stictoleptura, Strangalepta, Strangalia, Strophiona, Trachysida, 

Trigonarthris, Typocerus, Xestoleptura, Xylosteus. 

Cerambycinae Latreille 1802: Achryson, Aethecerinus, Agallissus, Amannus, Ancylocera, 

Aneflomorpha, Aneflus, Anelaphus, Anopliomorpha, Anoplocurius, Astromula, Atylostagma, 

Axestinus, Batyle, Brothylus, Cacophrissus, Callidiellum, Callidium, Callimoxys, Calloides, 

Callona, Chlorida, Chrotoma, Clytoleptus, Clytus, Coleomethia, Crioprosopus, Crossidius, 

Curius, Curtomerus, Cyrtophorus, Deltaspis, Dendrobias, Dicentrus, Dihammaphora, Dryobius, 

Eburia, Elaphidion, Elaphidionopsis, Elatotrypes, Elytroleptus, Enaphalodes, Eucrossus, 

Euderces, Eudistenia, Eumichthus, Euryscelis, Eustromula, Geropa, Glycobius, Gnaphalodes, 

35 

CLASSIFICATION

36 



Gracilia, Gymnopsyra, Haplidoeme, Haplidus, Hesperanoplium, Hesperophanes, Heterachthes, 

Heterops, Holopleura, Hybodera, Hylotrupes, Hypexilis, Knulliana, Lampropterus, Lianema, 

Linsleyonides, Lissonotus, Lophalia, Malacopterus, Malobidion, Mannophorus, Megacheuma, 

Megacyllene, Megaderus, Meganoplium, Megobrium, Meriellum, Metaleptus, Methia, 

Micraneflus, Micranoplium, Microclytus, Molorchus, Nathrius, Neaneflus, Neoclytus, Neocompsa, 

Nesostizocera, Obrium, Ochraethes, Odontocera, Oeme, Opsimus, Ornithia, Orwellion, Osmidus, 

Osmopleura, Paranoplium, Parelaphidion, Parevander, Penichroa, Pentanodes, Peranoplium, 

Perarthrus, Perigracilia, Phymatodes, Physocnemum, Piezocera, Placosternus, Plectromerus, 

Plesioclytus, Plinthocoelium, Plionoma, Poecilobrium, Pronocera, Pseudomethia, 

Pseudoperiboeum, Pseudopilema, Psyrassa, Purpuricenus, Rhodoleptus, Rhopalophora, 

Rhopalophorella, Romulus, Ropalopus, Rosalia, Sarosesthes, Schizax, Semanotus, Smodicum, 

Sphaerion, Stenaspis, Stenelaphus, Stenosphenus, Stromatium, Styloxus, Tanyochraethes, 

Taranomis, Tessaropa, Tetranodus, Tilloclytus, Tragidion, Triodoclytus, Tylcus, Tylonotus, 

Tylosis, Vandykea, Xeranoplium, Xylocrius, Xylotrechus, Zagymnus, Zamodes. 

Lamiinae Latreille 1825: Adetus, Aegomorphus, Alcidion, Alphomorphus, Amniscus, Astyleiopus, 

Astylidius, Ataxia, Cacostola, Callipogonius, Canonura, Cathetopteron, Ceratographis, 

Coenopoeus, Cyrtinus, Dectes, Desmiphora, Dorcaschema, Dorcasta, Ecyrus, Estoloides, 

Eupogonius, Eutrichillus, Glaucotes, Graphisurus, Goes, Hebestola, Hemierana, Hippopsis, 

Hyperplatys, Ipochus, Lagocheirus, Leptostylopsis, Leptostylus, Lepturges, Lochmaeocles, 

Lophopogonius, Lypsimena, Maculurges, Mecas, Microgoes, Moneilema, Monochamus, 

Neacanthocinus, Neoptychodes, Nyssodrysina, Oberea, Oncideres, Oplosia, Parmenonta, 

Parmenosoma, Peritapnia, Phaea, Plectrodera, Plectrura, Pogonocherus, Poliaenus, Psenocerus, 

Pseudastylopsis, Pygmaeopsis, Saperda, Spalacopsis, Steirastoma, Sternidius, Sternidocinus, 

Styloleptus, Synaphaeta, Tetraopes, Thryallis, Tigrinestola, Trichastylopsis, Trichocanonura, 

Tylocerina, Urgleptes, Valenus, Zaplous. 

Not placed in the database: Linsleyella. 

MEGALOPODIDAE Latreille 1802 

Zeugophorinae Böving and Craighead 1931: Zeugophora. 

ORSODACNIDAE Thomson 1859: 

Orsodacninae Thomson 1859: Orsodacne. 

Aulacoscelidinae Chapuis 1874: Aulacoscelis, Janbechynea. 

CHRYSOMELIDAE Latreille 1802 

Bruchinae Latreille 1802: Abutiloneus, Acanthoscelides, Algarobius, Althaeus, Amblycerus, 

Bruchidius, Bruchus, Callosobruchus, Caryedon, Caryoborus, Caryobruchus, Gibbobruchus, 

Kytorhinus, Megacerus, Meibomeus, Merobruchus, Mimosestes, Neltumius, Sennius, Stator, 

Stylantheus, Zabrotes. 

Donaciinae Kirby 1837: Donacia, Donaciella, Neohaemonia, Plateumaris, Poecilocera. 

Criocerinae Latreille 1804: Crioceris, Lema, Lilioceris, Neolema, Oulema. 

Hispinae Gyllenhal 1813: Agroiconota, Anisostena, Baliosus, Brachycoryna, Cassida, Chalepus, 

Charidiotella, Chelymorpha, Coptocycla, Deloyala, Erepsocassis, Eurypepla, Floridocassis, 

Glyphuroplata, Gratiana, Hemisphaerota, Hilarocassis, Jonthonota, Metrionella, 

Microctenochira, Microrhopala, Octotoma, Odontota, Opacinota, Parorectis, Pentispa, 

Physonota, Plagiometriona, Stenispa, Stenopodius, Strongylocassis, Sumitrosis, Xenochalepus. 

Chrysomelinae Latreille 1802: Calligrapha, Chrysolina, Chrysomela, Entomoscelis, 

Gastrophysa, Gonioctena, Hydrothassa, Labidomera, Leptinotarsa, Microtheca, Phaedon, 

Phratora, Plagiodera, Prasocuris, Timarcha, Zygogramma. 

Galerucinae Latreille 1802: Acalymma, Acrocyum, Agasicles, Agelastica, Altica, Amphelasma, 

Androlyperus, Aphthona, Argopistes, Asphaera, Blepharida, Brucita, Capraita, Cerotoma, 

Chaetocnema, Coraia, Crepidodera, Cyclotrypema, Derocrepis, Derospidea, Diabrotica, Dibolia, 

Disonycha, Distigmoptera, Dysphenges, Epitrix, Erynephala, Eusattodera, Galeruca, Galerucella, 

Glenidion, Glyptina, Hemiglyptus, Hippuriphila, Hornaltica, Hypolampsis, Keitheatus, 

Kuschelina, Longitarsus, Luperaltica, Luperosoma, Luperus, Lupraea, Lygistus, Lysathia, 

Macrohaltica, Malacorhinus, Mantura, Metrioidea, Miraces, Monocesta, Monomacra, Monoxia, 

Neobrotica, Omophoita, Ophraea, Ophraella, Orestioides, Orthaltica, Pachyonychus,


Paranapiacaba, Paratriarius, Phaedromus, Phrynocepha, Phydanis, Phyllecthris, Phyllobrotica, 

Phyllotreta, Pseudolampsis, Pseudoluperus, Pseudorthygia, Psylliodes, Pteleon, Pyrrhalta, 

Scelida, Scelolyperus, Sermylassa, Sphaeroderma, Strabala, Synetocephalus, Syphrea, Systena, 

Trachyscelida, Triarius, Trichaltica, Trirhabda. 

Lamprosomatinae Lacordaire 1848: Oomorphus. 

Cryptocephalinae Gyllenhal 1814: Anomoea, Babia, Bassareus, Chlamisus, Coleorozena, 

Coleothorpa, Coscinoptera, Cryptocephalus, Diachus, Diplacaspis, Exema, Griburius, 

Lexiphanes, Megalostomis, Neochlamisus, Pachybrachis, Pachyonychis, Pseudochlamys, Saxinis, 

Smaragdina, Triachus, Urodera. 

Eumolpinae Hope 1840: Brachypnoea, Bromius, Chrysochus, Colaspidea, Colaspis, Eumolpus, 

Euphrytus, Fidia, Glyptoscelis, Graphops, Megascelis, Metachroma, Metaparia, Metaxyonycha, 

Mychrous, Paria, Rhabdopterus, Spintherophyta, Syneta, Thricolema, Tymnes, Typophorus, 

Xanthonia. 

Superfamily CURCULIONOIDEA Latreille 1802 

NEMONYCHIDAE Bedel 1882 

Rhinorhynchinae Voss 1922: Atopomacer. 

Doydirhynchinae Pierce 1916: Cimberis, Lecontellus, Pityomacer. 

ANTHRIBIDAE Billberg 1820 

Anthribinae Billberg 1820: Allandrus, Anthribus, Araeoderes, Brevibarra, Discotenes, Eugonus, 

Euparius, Goniocloeus, Gymnognathus, Ischnocerus, Ormiscus, Phaenithon, Phoenicobiella, 

Piesocorynus, Stenocerus, Toxonotus, Trachytropis, Trigonorhinus, Tropideres. 

Choraginae Kirby 1819: Acaromimus, Araecerus, Choragus, Euxenulus, Euxenus, 

Pseudochoragus. 

BELIDAE Schönherr 1826 

Oxycoryninae Schönherr 1840: Rhopalotria. 

ATTELABIDAE Billberg 1820 

Rhynchitinae Gistel 1856: Auletobius, Deporaus, Eugnamptus, Haplorhynchites, Involvulus, 

Merhynchites, Pselaphorhynchites, Pterocolus, Rhynchites. 

Attelabinae Billberg 1820: Attelabus, Himatolobus, Homoeolabus, Xestolabus. 

BRENTIDAE Billberg 1820 

Cyladinae Schönherr 1823: Cylas. 

Brentinae Billberg 1820: Arrhenodes, Brentus, Heterobrentus, Paratrachelizus, Stereodermus. 

Nanophyinae Gistel 1856: Nanophyes. 

Apioninae Schönherr 1823: Apion, Chrysapion, Podapion. 

ITHYCERIDAE Schönherr 1823 

Ithycerus. 

CURCULIONIDAE Latreille 1802 

Brachycerinae Billberg 1820: Achrastenus, Acmaegenius, Adaleres, Agasphaerops, Agraphus, 

Agronus, Amotus, Anametis, Anchitelus, Aphrastus, Aracanthus, Aragnomus, Artipus, 

Atrichonotus, Barynotus, Barypeithes, Brachysomus, Brachystylus, Bradyrhynchoides, 

Callirhopalus, Calomycterus, Calyptillus, Cercopedius, Cercopeus, Chaetechidius, Cimbocera, 

Colecerus, Compsus, Crocidema, Cryptolepidus, Cyrtepistomus, Diamimus, Diaprepes, 

Dichoxenus, Dirotognathus, Dysticheus, Epicaerus, Ericydeus, Eucilinus, Eucyllus, Eudiagogus, 

Evotus, Geoderces, Geodercodes, Glaphyrometopus, Graphorhinus, Graphognathus, 

Hadromeropsis, Hormops, Hormorus, Hypera, Isodacrys, Isodrusus, Lepesoma, Lepidophorus, 

Leptopinara, Liophloeus, Listroderes, Listronotus, Lixellus, Lupinocolus, Melanolemma, 

Menoetius, Mesagroicus, Miloderes, Miloderoides, Minyomerus, Mitostylus, Myllocerus, 

Neoptochus, Omias, Omileus, Ophryastes, Orimodema, Orthoptochus, Otiorhynchus, Pachnaeus, 

37 

CLASSIFICATION

38 



Pachyrhinus, Pactorrhinus, Pandeleteinus, Pandeleteius, Panscopus, Pantomorus, 

Paracimbocera, Paragraphus, Paranametis, Paraptochus, Peritaxia, Peritelinus, Peritodes, 

Peritelopsis, Philopedon, Phyllobius, Phyxelis, Piscatopus, Plinthodes, Polydacrys, Polydrusus, 

Pseudocercopeus, Pseudocyphus, Pseudorimus, Rhigopsis, Rhypodillus, Sapotes, Scalaventer, 

Sciaphilus, Sciopithes, Sitona, Stamoderes, Stenoptochus, Stereogaster, Stomodes, Strophosoma, 

Tanymecus, Thecesternus, Thinoxenus, Thricolepis, Thricolepoides, Thylacites, Trachyphloeus, 

Trichalophus, Triglyphulus, Trigonoscuta, Trigonoscutoides, Tropiphorus, Vitavitus. 

Curculioninae Latreille 1802: Acalles, Acallodes, Acalyptus, Acanthoscelidius, Acentrinops, 

Acoptus, Alaocybites, Amalus, Ambates, Amercedes, Ampeloglypter, Anacentrinus, Anchonus, 

Anthonomopsis, Anthonomus, Apteromechus, Archarias, Auleutes, Aulobaris, Bagous, Baridius, 

Barilepis, Barilepton, Barinus, Baris, Brachybamus, Brachyogmus, Buchananius, Calandrinus, 

Calles, Campylirhynchus, Canistes, Catapastus, Centrinaspis, Centrinites, Centrinogyna, 

Centrinopus, Centrinus, Ceuthorhynchidius, Ceutorhynchus, Chalcodermus, Chaleponotus, 

Chelonychus, Chionanthobius, Cholinobaris, Cionomimus, Cionus, Cionopsis, Cleonis, 

Cnemogonus, Conotrachelus, Cophes, Copturus, Cosmobaris, Craponius, Cryptorhynchus, 

Curculio, Cylindridia, Cylindrocopturinus, Cylindrocopturus, Cyrtobagous, Desmoglyptus, 

Dietzella, Dietzianus, Diorymeropsis, Dirabius, Dorytomus, Eisonyx, Elleschus, Epacalles, 

Ephelops, Epimechus, Episcirrus, Erodiscus, Eubrychius, Eubulus, Eudociminus, Euhrychiopsis, 

Eulechriops, Eurhoptus, Euscepes, Faustinus, Gastrotaphrus, Geraeus, Gerstaeckeria, 

Gilbertiola, Glyptobaris, Gononotus, Grypus, Gymnetron, Hamaba, Haplostethops, Heilipus, 

Hesperobaris, Hohonus, Homorosoma, Hylobius, Hypocoeliodes, Hypoleschus, Hypurus, 

Idiostethus, Laemosaccus, Larinus, Lechriops, Lembodes, Lepyrus, Lignyodes, Liometophilus, 

Lixus, Lissorhoptrus, Lymantes, Macrorhoptus, Madarellus, Maemactes, Magdalinops, Magdalis, 

Mecinus, Metopotoma, Miarus, Micralcinus, Microbaris, Microcholus, Microhyus, Microlarinus, 

Micromastus, Mononychus, Myrmex, Nanops, Nanus, Narberdia, Nedyus, Neochetina, 

Neomastix, Neophycocoetes, Neophytobius, Neotylopterus, Neoulosomus, Nicentrus, Notaris, 

Notiodes, Notolomus, Ochyromera, Odontocorynus, Odontopus, Oligolochus, Onychobaris, 

Onychylis, Oomorphidius, Oopterinus, Orchidophilus, Orthoris, Pachybaris, Pachygeraeus, 

Pachylobius, Pachytychius, Paracamptus, Parenthis, Pelenosomus, Peltophorus, Peracalles, 

Perigaster, Pheloconus, Phyllotrox, Phyrdenus, Phrydiuchus, Phytobius, Piazorhinus, Pissodes, 

Plesiobaris, Plocamus, Plocetes, Pnigodes, Poophagus, Procas, Promecotarsus, 

Pseudanthonomus, Pseudoacalles, Pseudobaris, Pseudocentrinus, Pseudomopsis, Pseudomus, 

Psomus, Pycnobaris, Pycnogeraeus, Rhinocyllus, Rhinoncus, Rhoptobaris, Rhynchaenus, 

Rhynchus, Rhyssomatus, Rileyonymus, Rutidosoma, Schizomicrus, Sibariops, Sibinia, Sirocalodes, 

Smicraulax, Smicronyx, Stenobaris, Stenopelmus, Steremnius, Sternechus, Sternochetus, 

Stethobaris, Sthereus, Stictobaris, Strongylotes, Sudus, Tachygonus, Tachypterellus, Tanysphyrus, 

Trachodes, Tranes, Trepodaris, Trichobaris, Trichodirabius, Trichomagdalis, Trichosirocalus, 

Tychius, Tyloderma, Zascelis, Zygobarella, Zygobarinus, Zygobaris. 

Dryophthorinae Schönherr 1825: Cosmopolites, Dryophthorus, Metamasius, Orthognathus, 

Rhodobaenus, Rhynchophorus, Scyphophorus, Sitophilus, Sphenophorus, Yuccaborus. 

Cossoninae Schönherr 1825: Acamptus, Apotrepus, Caecossonus, Carphonotus, Caulophilus, 

Cossonus, Dryotribus, Dynatopechus, Elassoptes, Emphyastes, Eucoptus, Hexarthrum, Himatium, 

Macrancylus, Mesites, Micromimus, Nyssonotus, Pentarthrinus, Pentarthrum, Phloeophagus, 

Pselactus, Pseudopentarthrum, Rhamphocolus, Rhinanisus, Rhyncolus, Stenancylus, Stenomimus, 

Stenoscelis, Tomolips, Trichacorynus. 

Scolytidae Latreille 1804 

Hylesininae: Alniphagus, Carphobius, Carphoborus, Chaetophloeus, Chramesus, Cnesinus, 

Dendroctonus, Dendrosinus, Hylastes, Hylastinus, Hylesinus, Hylurgopinus, Hylurgops, 

Liparthrum, Phloeosinus, Phloeotribus, Plagiocerus, Polygraphus, Pseudohylesinus, Scierus, 

Xylechinus. 

Scolytinae Latreille 1804: Ambrosiodmus, Araptus, Cactopinus, Cnemonyx, Coccotrypes, 

Conophthorus, Corthylus, Cryphalus, Cryptocarenus, Crypturgus, Dendrocranulus, Dendroterus, 

Dolurgus, Dryocoetes, Ernoporicus, Gnathotrichus, Hylocurus, Hypocryphalus, Hypothenemus, 

Ips, Lymantor, Miracis, Micracisella, Monarthrum, Pityoborus, Pityogenes, Pityokteines, 

Pityophthorus, Pityotrichus, Onthotomicus, Premnobius, Procryphalus, Pseudopityophthorus,


Pseudothysanoes, Pycnarthrum, Scolytodes, Scolytogenes, Scolytus, Stenocleptus, Thysanoes, 

Trischidias, Trypodendron, Trypophloeus, Xyleborinus, Xyleborus, Xylosandrus, Xyloterinus. 

Platypodidae Shuckard 1840 

Platypus. 

39 

CLASSIFICATION

40 


NOMINA INSECTA NEARCTICA


COLEOPTERA 

41

42 

NOMINA INSECTA NEARCTICA

Classification; pp. 21 - Nearctica

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