Title of manuscript - Mycosphere-online journal

Mycosphere Doi 10.5943/mycosphere/3/2/5 

Towards incorporating anamorphic fungi in a natural classification – checklist 

and notes for 2011 

Wijayawardene DNN 1,2 , McKenzie EHC 3 and Hyde KD 1,2 

1 Institute of Excellence in Fungal Research, Mae Fah Luang University, Chiang Rai 57100, Thailand 

2 School of Science, Mae Fah Luang University, Chiang Rai 57100, Thailand 

3 Manaaki Whenua Landcare Research, Private Bag 92170, Auckland, New Zealand 

Wijayawardene DNN, McKenzie EHC, Hyde KD 2012 – Towards incorporating anamorphic fungi 

in a natural classification – checklist and notes for 2011. Mycosphere 3(2), 157-228, Doi 10.5943 

/mycosphere/3/2/5 

A complilation of anamorphic names for both Ascomycota and Basidiomycota is provided which 

comprises 2895 genera. The genera are listed against a backbone of teleomorphic relationships 

where known. The study reveals that 73 genera and 95 anamorph-like genera are linked to 

teleomorphic genera names, 447 genera (three anamorph-like genera) are linked to teleomorph 

families, orders or classes, while for more than 1592 (55%) genera no teleomorph link is known. 

The links are based on the literature and often because fungi were found in association, but have not 

been proven by molecular data. Many are based on links with species other than the generic type 

and thus must be considered questionable for the genus. A considerable effort is needed to establish 

whether these links are correct. 

Key words – Asexual fungi – life cycle – sexual fungi – taxonomy 

Article Information 

Received 24 March 2012 

Accepted 29 March 2012 

Published online 10 April 2012 

Corresponding author: KD Hyde – e-mail – kdhyde3@gmail.com 

Introduction 

The purpose of the present paper is to 

collate the knowledge of asexual fungi in 

relationship to their sexual states. It attempts to 

classify the asexual genera in a natural 

biological framework for the Ascomycota 

based on the Outline of Ascomycota 2009 

(Lumbsch & Huhndorf 2010) and for the 

Basidiomycota based on Kirk et al. (2008). The 

present paper has used data from the previous 

outline of Hyde et al. (2010) as well as 

literature published in 2011 and updated with 

opinions in Seifert et al. (2011). 

There is now a mandate in the systematics 

community to place one name on a 

holomorphic fungus species as opposed to the 

previous system (especially in Ascomycota) of 

naming both sexual and asexual morphs 

(Samuels et al. 2009, Hawksworth 2011; 

Hawksworth et al. 2011, Taylor 2011; Wingfield 

et al. 2011). Also, as more and more 

strains of asexual genera are sequenced 

(Shenoy et al. 2010, Põldmaa 2011, Reblova & 

Seifert 2011), these genera will be classified in 

families, orders and classes of Ascomycota and 

Basidiomycota. This will result in more and 

more asexual states being described and illustrated 

without being given formal names and 

therefore it is important to have a forum where 

asexual data can be brought to the attention of 

researchers. It is hoped that this compilation 

will facilitate the move towards one fungus one 

name (Shenoy et al. 2007, 2010, Hawksworth 

et al. 2011). By placing all asexual genera 

against the backbone classification of the sexual 

genera it will be possible to establish, 1) 

157

what are the sexual states of various species, 

genera,families or orders, 2) which genera have 

no information, 3) which are the older or more 

commonly used names for each genus, and 4) 

illustrate the redundancy of the dual classification 

system. 

This compilation also provides a single 

place where asexual connections can be accessed 

and critically scrutinized so that a stable and 

reliable system can be developed based on published 

data. We do not claim that this compilation 

is complete or definitive; however, it 

provides a forum for displaying data and can 

be added to or changed as more data becomes 

available or errors are discovered. In future 

issues we will explore various connections and 

provide more data. 

In the long term it will be desirable if 

Myconet (http://www.fieldmuseum.org/myconet) 

is merged with this document, however, 

this should occur at some time in the future, 

when more data is available. 

Material and methods 

The overall scheme for the classification 

of the sexual fungi follows the Outline of 

Ascomycota 2009 (Lumbsch & Huhndorf 

2010), with additional data on anamorphic 

Basidiomycota from Kirk et al. (2008). This 

scheme is chosen as the schemes in the 

Dictionary of the Fungi, Index Fungorum, and 

MycoBank differ to a greater or lesser extent 

and it was felt wise to follow what the authors 

consider is an admirable attempt to keep 

abreast of the current literarure as is routinely 

carried out in the Outline of Ascomycota. This 

compilation also annotates changes made in the 

form of notes on asexual fungi (marked with an 

asterisk after the entry, with notes in the later 

“Notes” section) so that changes can easily be 

followed; this is not always true of the 

available databases. 

Asexual genera or ‘-like’ genera are 

listed in bold under the appropriate fungal 

classification and the link to a teleomorph 

genus given where found. Dubious genera are 

excluded but can be added if they are later 

found to be good genera. Fossil fungi are also 

excluded as it would be impossible with 

present techniques available to provide a natural 

classification for them. A question mark 

before the entry means that its placement in 

158 


that family or order is not fully established and 

generally follows Lumbsch & Huhndorf 

(2010). A question mark before anamorph 

means it is not clear that this is the anamorph 

of this genus, although there are many cases 

without a question mark where the link has not 

equivocally been proven. An asterisk following 

the names indicates an entry in the “Notes” 

section. 

In many genus records in Index Fungorum 

the species is linked to more than one 

anamorphic genus. Such records can be located 

by searching the genus throughout the list for 

repeat entries. The synonyms of genera are not 

listed as they can be easily found in Index 

Fungorum. 

We encourage colleagues to inform us of 

any missing entries, new data or misunderstandings, 

which will be updated in future 

compilations. We would also invite experts of 

specific groups to contribute to future compilations 

on their specific group(s) of interest and 

encourage reviews or illustrated accounts of 

families, orders or other groups relating to 

sexual and asexual links. Reviews of asexual 

genera with sections or series, such as 

Aspergillus, Penicillium and Verticillium, the 

powdery mildews, many of the yeasts and 

yeast-like genera and the asexual smuts and 

rusts are particularly needed. Similarly the 

lichenized ascomycete asexual states are poorly 

documented. 

As there is mandate towards using one 

name for both the sexual and asexual states of a 

fungus we also encourage authors to submit 

articles or opinions concerning the use of either 

name. This might be in the form of a general 

critic, arguments for or against using one name, 

or an article that specifically addresses an 

individual group or genus. 

In the present outline we have added 

(H) = hyphomycete, (C) = coelomycete, (Y) = 

yeast and (R) = Rust against entries where 

known. Although this grouping is artificial we 

do this to facilitate finding data on either as the 

groups are generally dealt with in different text. 

For example, data for the hyphomycetes have 

recently been complied and updated by Seifert 

et al. (2011) and the new data for hyphomycetes 

included here have been extracted from this 

tome. Data for coelomycetes were extracted 

from Sutton (1980), Nag Raj (1993), http://

www. Cybertr uffle.org.uk/cybernome/eng/index.htm 

(Accesion date 18.03.2011) and http://www.landcarer 

esearch .co.nz/html /data asp? 

ID = &NAMEKey = 26885 (Accesion date 

6.12.2011). 

Index to Ascomycota 

Phylum ASCOMYCOTA Caval-Sm. 

Subphylum TAPHRINOMYCOTINA O.E. 

Erikss. & Winka 

Class Taphrinomycetes O.E. Erikss. & Winka 

Taphrinales Gäum. & C.W. Dodge 

Protomycetaceae Gray 

Saitoella Goto, Sugiy., Hamam. & Komag. 

1987 (Y) 

Taphrinaceae Gäum. 

Lalaria R.T. Moore 1990, anamorphic 

Taphrina Fr. 1815 (Y) 

Subphylum SACHAROMYCOTINA O.E. 


Class Saccharomycetes O.E. Erikss. & Winka 

Saccharomycetales Kudrjanzev 

Dipodascaceae Engl. & E. Gilg 

Blastobotrys Klopotek 1967, anamorphic 

Trichomonascus H.S. Jacks. 1948 (H) 

?Candida Berkhout 1923, anamorphic 

Sporopachydermia Rodr. Mir. 1978 (Y) 

Geotrichum Link 1809, anamorphic 

Galactomyces Redhead & Malloch 

1977 and Dipodascus Lagerh. 1892 (H) 

Geotrichum-like, anamorphic Basidioascus 

Matsush. 2003 (H) 

Saprochaete Coker & Shanor 1939, anamorphic 

Magnusiomyces Zender 1926 

(H) 

Endomycetaceae J. Schröt. 

Fusidium Link 1809, possibly anamorphic 

Ascocephalophora K. Matsush. & 


Lipomycetaceae E.K. Novák & Zsolt 

Myxozyma Van der Walt, Weijman & Arx 

1981 (Y) 

Metschnikowiaceae T. Kamienski 

Candida Berkhout 1923, anamorphic 

Clavispora Rodr. Mir. 1979 and 


Metschnikowia T. Kamienski 1899 (Y) 

Pichiaceae Zender 

Brettanomyces N.H. Claussen ex Custers 

1940, anamorphic Dekkera Van der 

Walt 1964 (Y) 

Eeniella M.T. Sm., Bat. Vegte & Scheffers 

1981, anamorphic Dekkera Van der 

Walt (Y) 

Enantiothamnus Pinoy 1911 (Y) 

Hyphopichia Arx & Van der Walt 1976 

(Y) 

Saccharomycetaceae G. Winter 


Ogataea Y. Yamada, K. Maeda & 

Mikata 1994, Lodderomyces Van der 

Walt 1966 and Spathaspora N.H. 

Nguyen, S.O. Suh & M. Blackw. 2006 

(Y) 

Saccharomycodaceae Kudrjanzev 


Lodderomyces Van der Walt 1966 and 

Spathaspora N.H. Nguyen, S.O. Suh & 

M. Blackw. 2006 (Y) 

Kloeckera Janke 1923, anamorphic Hanseniaspora 

Zikes 1911 (Y) 

Trichomonascaceae Kurtzman & Robnett 


Zygoascus M.T. Sm. 1986 (Y) 

Saccharomycetales, genera incertae sedis 

Aciculoconidium D.S. King & S.C. Jong 

1976 (Y) 

Botryozyma Shann & M.T. Sm. 1992, 

anamorphic Ascobotryozyma J. Kerrigan, 

M.T. Sm. & J.D. Rogers 2001 (Y) 


Starmerella C.A. Rosa & Lachance 

1998 (Y) 

Cicadomyces Šulc 1911 (Y) 

Macrorhabdus Tomasz., Logan, Snowden, 

Kurtzman & Phalen 2003 (Y) 

Oosporidium Stautz 1931 (Y) 

Pseudomycoderma H. Will 1916 (Y) 

Schizoblastosporion Cif. 1930 (Y) 

Trigonopsis Schachner 1929 (Y) 

Saccharomycetes, genera incertae sedis 

Selenotila Lagerh. 1892 (Y) 

159

160 

Selenozyma Yarrow 1977 (Y) 

Subphylum PEZIZOMYCOTINA O.E. 


Class Arthoniomycetes O.E. Erikss. & Winka 

Arthoniales Henssen ex D. Hawksw. & O.E. 

Erikss. 

Arthoniaceae Reichenb. ex Reichenb. 

Helicobolomyces Matzer 1995, anamorphic 

Arthonia Ach. 1806 (C) 

?Septocyta Petr. 1927, anamorphic Arthonia 

Ach. 1806 (C) 

Subhysteropycnis Wedin & Hafellner 

1998, anamorphic Arthonia Ach. 1806 

(C) 

Roccellaceae Chevall. 

Sporhaplus H.B.P. Upadhyay 1964, anamorphic 

Mazosia A. Massal 1854 (C) 

Class Dothideomycetes sensu O.E. Erikss. & 

Winka 

Subclass Dothideomycetidae P.M. Kirk, P.F. 

Cannon, J.C. David & J.A. Stalpers ex C.L. 

Schoch, Spatafora, Crous & Shoemaker 

Capnodiales Woron. 

Antennulariellaceae Woron. 

Capnodendron S. Hughes 1976, anamorphic 

Antennulariella Woron. 1915 (H) 

Capnofrasera S. Hughes 2003 (H) 

Heteroconium Petr. 1949, anamorphic 

Antennulariella Woron. 1915 (H) 

Capnodiaceae (Sacc.) Höhn. ex Theiss. 

Acanthorus Bat. & Cavalc. 1967 (C) 

?Antennariella Bat. & Cif. 1963 (C)* 

Apiosporium Kunze 1817 (C) 

Conidiocarpus Woron. 1927, anamorphic 

Phragmocapnias Theiss. & Syd. 1918 

and Scorias Fr. 1825 (C)* 

Fumagospora G. Arnaud 1911, anamorphic 

Capnodium Mont. 1848 (C) 

?Fumiglobus D.R. Reynolds & G.S. 

Gilbert 2006 (C)* 

Microxiphium (Harv. ex Berk. & Desm.) 

Thüm. 1879 (as Microxyphium) (H)* 

?Mycogelidium W.Y. Zhuang 2007 (C) 

Phaeoxyphiella Bat. & Cif. 1963, anamorphic 

Capnodium Mont. 1848 (C)* 

?Polychaetella Speg. 1918, anamorphic 

Capnodium Mont. 1848 (C)* 

Polychaeton (Pers.) Lev. 1846 (C)* 


Scolecoxyphium Cif. & Bat. 1956, 

anamorphic Capnodium Mont. 1848 

and Scorias Fr. 1825 (C)* 

Tripospermum Speg. 1918 anamorphic 

Triposporiopsis W.Yamam 1955 (H)* 

Coccodiniaceae Höhn. ex O.E. Erikss. 

Bisbyopeltis Bat. & A.F. Vital 1957 (C) 


Thüm. 1879, anamorphic Dennisiella 

Bat. & Cif. 1962 (H)* 

Davidiellaceae C.L. Schoch, Spatafora, Crous 

& Shoemaker 

Acroconidiella J.C. Lindq. & Alippi 1964, 

anamorphic Davidiella Crous & U. 

Braun 2003 (H) 

Cladosporium Link 1816, anamorphic 

Davidiella Crous & U. Braun 2003 (H) 

Dichocladosporium K. Schub., U. Braun 

& Crous 2007 (C) 

Graphiopsis Trail 1889, anamorphic 

Davidiella Crous & U. Braun 2003 (H) 

Hoornsmania Crous 2007 (H) 

Rachicladosporium Crous, U. Braun & 

C.F. Hill 2007 (H) 

Toxicocladosporium Crous & U. Braun 

2007 (H) 

Verrucocladosporium K. Schub., Aptroot 

& Crous 2007 (H) 

Dissoconiaceae Crous & de Hoog 

Dissoconium de Hoog, Oorschot & Hijwegen 

1983 (H) 

Ramichloridium Stahel 1937 ex de Hoog 

1977 (H) 

Metacapnodiaceae S. Hughes & Corlett 

Capnobotrys S. Hughes 1970, anamorphic 

Metacapnodium Speg. 1918 (H) 

Capnocybe S. Hughes 1966, anamorphic 


Capnophialophora S. Hughes 1966, anamorphic 

Metacapnodium Speg. 1918 

(H) 

Capnosporium S. Hughes 1976, anamorphic 


Hormiokrypsis Bat. & Nascim. 1957, anamorphic 

Metacapnodium Speg. 1918 

and Ophiocapnocoma Bat. & Cif. 1963 

(H) 

Torulopsiella Bender 1932, anamorphic

Ophiocapnocoma Bat. & Cif. 1963 and 


Mycosphaerellaceae Lindau 

Anguillosporella U. Braun 1995 (H) 

Asperisporium Maubl. 1913 (H) 

Asteromella-like, anamorphic Gillotia 

Sacc. & Trotter 1913 (C) 

Cercospora Fresen. 1863, anamorphic 

Mycosphaerella Johanson 1884, Sphaerulina 

Sacc. and others (H) 

Cercosporella Sacc. 1880, anamorphic 

Mycosphaerella Johanson 1884 (H) 

?Clypeispora A.W. Ramaley 1991, anamorphic 

Mycosphaerella Johanson 

1884 (C) 

Colletogloeum Petr. 1953, anamorphic 

Mycosphaerella Johanson 1884 (C) 

Coniothryium-like, anamorphic Bruneosphaerella 

Crous 2009 (C) 

Deightoniella S. Hughes 1952 (H) 

Didymochora Höhn. 1918, anamorphic 

Euryachora Fuckel 1870 (C) 

Distocercospora N. Pons & B. Sutton 

1988 (H) 

Dothistroma Hulbary 1941 (C) 

Floricola Kohlm. & Volkm.Kohlm. 2000 

(C) 

Fusicladiella Höhn. 1919, anamorphic 


Laocoön J.C. David 1997 (H) 

Lecanosticta Syd. 1922 (C) 

Microcyclospora Jana Frank, Schroers & 

Crous 2010 (H) 

Microcyclosporella Jana Frank, Schroers 

& Crous 2010 (H) 

Miuraea Hara 1948, anamorphic Mycosphaerella 

Johanson 1884 (H) 

Mycopappus Redhead & G.P. White 1985, 

anamorphic Mycosphaerella Johanson 

1884 (H) 

Mycovellosiella Rangel 1918, anamorphic 


Parastenella J.C. David 1991 (H) 

Passalora Fr. 1849, anamorphic Mycosphaerella 


Penidiella Crous & U. Braun 2007 (H) 

Periconiella Sacc. 1885, anamorphic 

Allosoma Syd. 1926 (H) 

Phacellium Bonord. 1860, anamorphic 


Phaeoisariopsis Ferraris 1909, anamorphic 



Phaeophleospora Rangel 1916 (C) 

Phaeothecoidea Crous 2007 (H) 

Phloeospora Wallr. 1833, anamorphic 

Mycosphaerella Johanson 1884 (C) 

Placosphaeria-like, anamorphic Euryachora 

Fuckel 1870 (C) 

Polythrincium Kunze 1817, anamorphic 

Cymadothea F.A. Wolf 1935 (H) 

Pseudocercospora Speg. 1910 anamorphic 


Pseudocercosporella Deighton 1973, anamorphic 

Mycosphaerella Johanson 

1884 (H) 

Pseudocercosporidium Deighton 1973 (H) 

Quasiphloeospora B. Sutton, Crous & 

Shamoun 1996 (H) 

Ramichloridium Stahel ex de Hoog 1977 

(H) 

Ramichloridium-like, anamorphic Mycosphaerella 


Ramularia Unger 1833, anamorphic Mycosphaerella 

Johanson 1884 and 

Melanodothis R.H. Arnold 1972 (H) 

Ramulariopsis Speg. 1910, anamorphic 


Ramulispora Miura 1920, anamorphic 


Rhabdospora (Durieu & Mont. ex Sacc.) 

Sacc. 1884 (C) 

Rhexocercosporidium U. Braun 1994 (H) 

Semipseudocercospora J.M. Yen 1983 (H) 

Septoria Sacc. 1884 (C) 

Septoria-like, anamorphic Mycosphaerella 

Johanson 1884 (C) 

Sirosporium Bubák & Serebrian. 1912 (H) 

Sonderhenia H.J. Swart & J. Walker 1988 

(C) 

Stenella Syd. 1930, anamorphic Mycosphaerella 


Stenellopsis B. Huguenin 1966 (H) 

Stigmina Sacc. 1980, anamorphic Mycosphaerella 


Thedgonia B. Sutton 1973, anamorphic 


Thedgonia-like (H) 

Trochophora R.T. Moore 1955 (H) 

Verrucisporota D.E. Shaw & Alcorn 1993 

(H) 

Xenostigmina Crous 1998, anamorphic 

Mycosphaerella Johanson 1884 or 

Didymella Sacc. ex D. Sacc. 1880 (H) 

161

162 

Zasmidium Fr. 1849, anamorphic Mycosphaerella-like 

and Rasutoria M.E. Barr 

1987 (H) 

Zymoseptoria Quaedvlieg & Crous 2011 

(C)* 

Capnodiales, genera incertae sedis 

Capnocheirides J.L. Crane & S. Hughes 

1982 (H) 

Heptaster Cif., Bat. & Nascim. 1956 (H) 

Hormonema-like, anamorphic Xenomeris 

Syd. 1924 (H) 

Houjia G.Y. Sun & Crous 2010 (H) 

Micropustulomyces R.W. Barreto 1995 (C) 

Phaeotheca Sigler, Tsuneda & J.W. 

Carmich. 1981 (H) 

Phaeothecoidiella Batzer & Crous 2010 

(H) 

Pseudovirgaria H.D. Shin, U. Braun, 

Arzanlou & Crous 2007 (H) 

Rachicladosporium Crous, U. Braun & 

C.F. Hill 2007 (H) 

Recurvomyces Selbmann & de Hoog 2008 

(H) 

Scolecostigmina U. Braun 1999 (H) 

Sporidesmajora Batzer & Crous 2010 (H) 

Verrucocladosporium K. Schub., Aptroot 

& Crous 2007 (H) 

Dothideales Lindau 

Dothideaceae Chevall. 

Lecanosticta Syd. 1922, anamorphic Scirrhia 

Nitschke ex Fuckel 1870 (C) 

Podoplaconema Petr. 1921, anamorphic 

Omphalospora Theiss. & Syd. 1915 (C) 

Dothioraceae Theiss. & H. Syd. 

Aureobasidium Viala & G. Boyer 1891, 

anamorphic Columnosphaeria Munk 

1953 and Saccothecium Fr. 1836 (H) 

Dothichiza Lib. ex Roum. 1880, anamorphic 

Dothiora Fr. 1849 and Sydowia 

Bres. 1895 (C) 

Hormonema Lagerb. & Melin 1927, anamorphic 

Dothiora Fr. 1849, Pringsheimia 

Schulzer 1866 and Sydowia 

Bres. 1895 (H) 

Japonia Höhn. 1909, anamorphic Yoshinagaia 

Henn. 1904 (C) 

Kabatina R. Schneid. & Arx 1966 (C) 

?Rhizosphaera L. Mangin & Har. 1907, 

anamorphic Phaeocryptopus Naumov 


1915 and Dimerina Theiss. 1912 (C) 

Sclerophoma Höhn. 1909, anamorphic 

Sydowia Bres. 1895 (C) 

Teratosphaeriaceae Crous & U. Braun 

Acidomyces B.J. Baker, M.A. Lutz, S.C. 

Dawson, P.L. Bond & Banfield ex 

Selbmann, de Hoog & De Leo 2008 (H) 

Batcheloromyces Marasas, P.S. van Wyk 

& Knox-Dav. 1975 (H) 

Baudoinia J.A. Scott & Unter. 2007 (H) 

Capnobotryella Sugiy. 1987 (H) 

Catenulostroma Crous & U. Braun 2007, 

anamorphic Teratosphaeria Syd. & P. 

Syd. 1912 (H) 

Cibiessia Crous 2007 (H) 

Colletogloeopsis Crous & M.J. Wingf. 

1997, anamorphic Teratosphaeria Syd. 

& P. Syd. 1912 (C) 

Davisoniella H.J. Swart 1988 (C) 

Devriesia Seifert & N.L. Nick. 2004 (H) 

Elasticomyces Zucconi & Selbmann 2008 

(H) 

Friedmanniomyces Onofri 1999 (H) 

Hispidoconidioma Tsuneda & Davey 2010 

(H) 

Hobsoniopsis D. Hawksw. 2001 (H) 

Hortaea Nishim. & Miyaji 1984 (H) 

Kirramyces J. Walker, B. Sutton & Pascoe 

1992, anamorphic Teratosphaeria Syd. 

& P. Syd. 1912 (C) 

Nothostrasseria Nag Raj 1983 (C) 

Passalora Fr. 1849 (H) 

Penidiella Crous & U. Braun 2007 (H) 

Phacellium Bonord. 1860 (H) 

Phaeothecoidea Crous 2007 (H) 

Pseudocercosporella Deighton 1973 (H) 

Pseudoramichloridium Cheewangkoon & 


Pseudotaeniolina J.L. Crane & Schokn. 

1986 (H) 

Ramichloridium Stahel ex de Hoog 1977 

(H) 

Recurvomyces Selbmann & de Hoog 2008 

(H) 

Readeriella Syd. & P. Syd. 1908 (C) 

Sporidesmium Link 1809 (H) 

Staninwardia B. Sutton 1971 (C) 

Stenella Syd. 1930 (H) 

Trimmatostroma-like, anamorphic Teratosphaeria 

Syd. & P. Syd. 1912 (H) 

Tripospermum Speg. 1918 (H)

Myriangiales Starbäck 

Elsinoaceae Höhn. ex Sacc. & Trotter 

Endosporium Tsuneda 2008 (H) 

Sphaceloma de Bary 1874 anamorphic 

Elsinoë Racib. 1900 (C) 

Xenodiella Syd. 1935, anamorphic Xenodium 

Syd. 1935 (H) 

Myriangiales, genera incertae sedis 

Phaeosclera Sigler, Tsuneda & J.W. Carmich. 

1981 (H) 

Sarcinomyces Lindner 1898 (H) 

Zopfiaceae G. Arnaud ex D. Hawksw. 1992 

Cladosporium-like, anamorphic Zopfia Rabenh. 

1874 (H) 

Subclass Pleosporomycetidae C.L. Schoch, 

Spatafora, Crous & Shoemaker 

Pleosporales Luttrell ex M.E. Barr 

Amniculicolaceae Yin. Zhang, C.L. Schoch, J. 

Fourn., Crous & K.D. Hyde 

Anguillospora Ingold 1942 (H) 

Spirosphaera Beverw. 1953 (H) 

Cucurbitariaceae G. Winter 

Camarosporium Schulzer 1870, anamorphic 

Cucurbitaria Gray 1821 (C) 

Coniothyrium-like, anamorphic Curreya 

Sacc. 1883 (C) 

Diplodia-like, anamorphic Cucurbitaria 

Gray 1821 (C) 

?Megaloseptoria Naumov 1925, anamorphic 


?Phaeoseptoria Speg. 1908, anamorphic 

Rhytidiella Zalasky 1968 (C) 

Phialophorophoma Linder 1944 (C) 

Phoma Sacc. 1880 (C) 

Pleurophoma Höhn. 1914 (C) 

Pleurostromella Petr. 1922, anamorphic 

Cucurbitaria Gray 1821 and Gibberidea 


Pyrenochaeta De Not. 1849, anamorphic 


Pyrenochaetopsis Gruyter, Aveskamp, & 

Verkley, 2010 (C) 

Syntholus A.W. Ramaley & M.E. Barr 

1997, anamorphic Syncarpella Theiss. 

& Syd. 1915 (C) 

Diademaceae Shoemaker & C.E. Babc. 

Alternaria-like, anamorphic Comoclathris 


Clem. 1909 (H)* 

Fusicladiella Höhn. 1919, anamorphic 

Didymosphaeria Fuckel 1870 (H) 

Didymellaceae Gruyter, Aveskamp & Verkley 

Ascochyta Lib. 1830, anamorphic Didymella 

Sacc. ex D. Sacc. 1880 (C) 

Ampelomyces Ces. ex Schltdl. 1852 (C) 

Boeremia Aveskamp, Gruyter & Verkley 

2010 (C) 

Chaetasbolisia Speg. 1918 (C) 

Dactuliochaeta G.L. Hartm. & J.B. Sinclair 

1988, anamorphic Didymella Sacc. 

1880 (C) 

Microsphaeropsis Höhn. 1917 (C) 

Peyronellaea Gold. ex Togliani 1952 (C) 

Phoma Sacc. 1880, anamorphic Didymella 

Sacc. ex D. Sacc. 1880 (C) 

Piggotia Berk. & Broome 1851, possibly 

anamorphic Platychora Petr. 1925 (C) 

Pithomyces Berk. & Broome 1875, anamorphic 

Leptosphaerulina McAlpine 

1902 (H) 

Stagonosporopsis Died. 1912, anamorphic 

Didymella Sacc. ex D. Sacc. 1880 (C) 

(=Ascochyta Lib. 1830) 

Didymosphaeriaceae Munk 

Cytoplea Bizz. & Sacc. 1885, anamorphic 

Roussoëlla Sacc. 1888 (C) 

Dothidotthiaceae Crous & A.J.L. Phillips 

?Dothiorella Sacc. 1880, anamorphic Dothidotthia 

Höhn. 1918 (C) 

Thyrostroma Höhn. 1911, anamorphic 

Dothidotthia Höhn. 1918 (H) 

Hypsostromataceae Huhndorf 

Pleurophomopsis-like, anamorphic Hypsostroma 

Huhndorf (C) 

Lentitheciaceae Yin. Zhang, C.L. Schoch, J. 

Fourn., Crous & K.D. Hyde 

Stagonospora (Sacc.) Sacc. 1884 (C) 

Leptosphaeriaceae M.E. Barr 

Camposporium Harkn. 1884, anamorphic 

Leptosphaeria Ces. & De Not. 1863 

(H) 

Coniothyrium Corda 1840, anamorphic 

Leptosphaeria Ces. & De Not. 1863 (C) 

Coniothyrium-like, anamorphic Neophaeo- 

163

164 

sphaeria M.P.S. Câmara, M.E. Palm & 

A.W. Ramaley 2003 (C) 

Phoma Sacc. 1880, anamorphic Leptosphaeria 

Ces. & De Not. 1863 (C) 

Plenodomus Preuss 1851 (C) 

Pyrenochaeta De Not. 1849, possibly 

anamorphic Leptosphaeria Ces. & De 

Not. 1863 (C) 

Lindgomycetaceae K. Hiray., Kaz. Tanaka & 

Shearer 

Taeniolella S. Hughes 1958 (H) 

Lolia Abdel-Aziz & Abdel-Whab 2010 

(C)* 

Lophiostomataceae Sacc. 

Anguillospora Ingold 1942, anamorphic 

Massarina Sacc. 1883 (H) 

Ascochyta Lib. 1830, anamorphic Trichometasphaeria 

Munk 1953 (C) 

Periconia Tode 1791 anamorphic Lophiostoma 

Ces. & De Not. 1863 (H) 

Pleuorphomopsis-like, anamorphic Lophiostoma 

Ces. & De Not. 1863 (C) 

Pyricularia Sacc. 1880, anamorphic Massarina 

Sacc. 1883 (H) 

Tumularia Descals & Marvanová 1987(H) 

Massariaceae Nitschke 

?Aplosporella-like, anamorphic Dubitatio 

Speg. 1882 (C) 

Myxocyclus Riess 1852, anamorphic Massaria 

De Not. 1844 (C) 

Torula Pers. 1794 (H) 

Massarinaceae Munk 

Aquaticheirospora Kodsueb & W.H. Ho 

2007 (H) 

Ceratophoma Höhn. 1917, anamorphic 

Massarina Sacc. 1883 (C) 

Cheirosporium L. Cai & K.D. Hyde 2008 

(H) 

Clavariopsis De Wild. 1895 (H) 

Helminthosporium Link 1809, anamorphic 

Splanchnonema Corda 1829 (H) 

Neottiosporina Subram. 1961 (C) 

Pseudodictyosporium Matsush. 1971 (H) 

Melanommataceae G. Winter 

Aposphaeria Sacc. 1880 (C) 

Beverwykella Tubaki 1975 (H) 

Exosporiella P. Karst. 1892, anamorphic 


Anomalemma Sivan. 1983 (H) 

Nigrolentilocus R.F. Castañeda & Heredia 

2001, anamorphic Melanomma Nitschke 

ex Fuckel 1870 (H) 

Monodictys S. Hughes 1958, anamorphic 

Ohleria Fuckel 1868 (H) 

Monotosporella S. Hughes 1958 (H) 

Pyrenochaeta De Not. 1849, anamorphic 

Byssosphaeria Cooke 1879 (C) 

Sporidesmiella P.M. Kirk 1982 (H) 

Montagnulaceae M.E. Barr 

Aschersonia Mont. 1848, anamorphic 

Montagnula Berl. 1896 (C) 

Microdiplodia Allesch. 1901, possibly 

anamorphic Karstenula Speg. 1879 (C) 

Microsphaeropsis Höhn. 1917, anamorphic 

Paraphaeosphaeria O.E. Erikss. 

1967 (C) 

Paraconiothyrium Verkley 2004, possibly 

anamorphic Paraphaeosphaeria O.E. 

Erikss. 1967 (C) 

Phaeosphaeriaceae M.E. Barr 

Amarenographium O.E. Erikss. 1982, anamorphic 

Amarenomyces O.E. Erikss. 

1981 (recently synonymised with 

Phaeosphaeria I. Miyake 1909) (C) 

Ampelomyces Ces. ex Schltdl. 1852 (C) 

Chaetosphaeronema Moesz 1915 (C) 

Coniothyrium Corda 1840, anamorphic 

Ophiobolus Riess 1854 (C) 

Harpophora W. Gams 2000, anamorphic 

anamorphic Ophiobolus Riess 1854 (H) 

Hendersonia-like, anamorphic Austropleospora 

R.G. Shivas & L. Morin 2010 

and Phaeosphaeria I. Miyake 1909 (C) 

Mauginiella Cavara 1925 (H) 

Neosetophoma Gruyter, Aveskamp & 

Verkley 2010 (C) 

?Parahendersonia A.W. Ramaley 1995, 

anamorphic Chaetoplea (Sacc.) Clem. 

1931 (C) 

Paraphoma Morgan-Jones & J.F. White 

1983 (C) 

Phaeoseptoria Speg. 1908, anamorphic 

Phaeosphaeria I. Miyake 1909 (C) 

Phaeostagonospora A.W. Ramaley 1997, 

anamorphic Phaeosphaeriopsis M.P.S. 

Câmara, M.E. Palm & A.W. Ramaley 

2003 (C) 

Phoma Sacc. 1880, anamorphic Ophio-

olus Riess 1854 (C) 

Rhabdospora (Durieu & Mont. ex Sacc.) 

Sacc. 1884, anamorphic Ophiobolus 

Riess 1854 (C) 

Scolecosporiella Petr. 1921, anamorphic 

Ophiosphaerella Speg. and Phaeosphaeria 

I. Miyake 1909 (C) 

Setophoma Gruyter, Aveskamp & Verkley 

2010 (C) 

?Sphaerellopsis Cooke 1883, anamorphic 

Eudarluca Speg. 1908 (C) 

Stagonospora (Sacc.) Sacc. 1884, anamorphic 

Phaeosphaeria I. Miyake 1909 (C) 

Tiarospora Sacc. & Marchal 1885 (C) 

Pleomassariaceae M.E. Barr 

Ceuthodiplospora Died. 1912, anamorphic 

Splanchnonema Corda (C) 

Corynespora Güssow 1906, anamorphic 

Pleomassaria Speg. 1880 (H) 

Myxocyclus Riess 1852, anamorphic 

Splanchnonema Corda 1829 (C) 

?Prosthemium Kunze 1817, anamorphic 

Pleomassaria Speg. 1880 (C) 

Scolicosporium Lib. ex Roum. 1880, anamorphic 

Asteromassaria Höhn. 1917 

(C) 

Shearia Petr. 1924, anamorphic Pleomassaria 

Speg. 1880 (C) 

Stegonsporium Corda 1827, anamorphic 

Splanchnonema Corda 1829 (C) 

Pleosporaceae Nitschke 

Alternaria Nees 1816, anamorphic Lewia 

M.E. Barr & E.G. Simmons 1986 (H)* 

Alternaria-like anamorphic Clathrospora 

Rabenh. 1857 (H) 

Alternariaster E.G. Simmons 2007 (H) 

Brachycladium Corda 1838, anamorphic 

Crivellia Shoemaker & Inderbitzin 

2006 (H) 

Bipolaris Shoemaker 1959, anamorphic 

Cochliobolus Drechsler 193 (H) 

Chalastospora E.G. Simmons 2007 (H) 


Pyrenophora Fr. 1849 (H) 

Curvularia Boedijn 1933, anamorphic 

Cochliobolus Drechsler 1934 (H) 

Dactuliophora C.L. Leakey 1964 (H) 

Dendryphiella Bubák & Ranoj. 1914, 

anamorphic Pleospora Rabenh. ex Ces. 

& De Not. 1863 (H) 


Dendryphion Wallr. 1833, anamorphic 

Pleospora Rabenh. ex Ces. & De Not. 

1863 (H) 

Dictyosporium Corda 1836 (H) 

Digitodesmium P.M. Kirk 1981 (H) 

Drechslera S. Ito 1930, anamorphic Pyrenophora 

Fr. 1849 (H) 

Edenia M.C. González, Anaya, Glenn, 

Saucedo & Hanlin 2007 (H) 

Embellisia E.G. Simmons 1971, anamorphic 

Lewia M.E. Barr & E.G. Simmons 

(H) 

Exserohilum K.J. Leonard & Suggs 1974, 

anamorphic Setosphaeria K.J. Leonard 

& Suggs 1974 (H) 

Gibbago E.G. Simmons 1986 (H) 

Lemonniera De Wild. 1894 (H) 

Marielliottia Shoemaker 1999, anamorphic 

Pyrenophora Fr. 1849 (H) 

Nimbya E.G. Simmons 1989, anamorphic 

Macrospora Fuckel 1870 (H) 

Prathoda Subram. 1956 (H) 

Sinomyces Yong Wang bis & X.G. Zhang 

2011 (H) 

Stemphylium Wallr. 1833, anamorphic Pleospora 

Rabenh. ex Ces. & De Not. 

1863 (H) 

Teretispora E.G. Simmons 2007 (H) 

Ulocladium Preuss 1851, anamorphic Lasiobotrys 

Kunze 1823 (H) 

Undifilum Pryor, Creamer, Shoemaker, 

McLain-Romero & Hambl. 2009 (H) 

Sporormiaceae Munk 

?Amorosia Mantle & D. Hawksw. 2006 

(H) 


Phoma-like, anamorphic Westerdykella 

Stolk 1955 (C) 

Teichosporaceae M.E. Barr 

Coniothyrium-like, anamorphic Immothia 

M.E. Barr (C) 

Tetraplosphaeriaceae Kaz. Tanaka & K. Hiray 

Pseudotetraploa Kaz. Tanaka & K. 

Hirayama 2009 (H) 

Quadricrura Kaz. Tanaka, K. Hirayama & 

Sat. Hatak. 2009 (H) 

Tetraploa Berk. & Broome 1850, anamorphic 

Tetraplosphaeria Kaz. Tanaka 

& K. Hirayama 2009 (H) 

165

166 

Tetraploa-like, anamorphic Triplosphaeria 

Kaz. Tanaka & K. Hirayama 2009 and 

Polyplosphaeria Kaz. Tanaka & K. 

Hirayama 2009 (H) 

Trematosphaeriaceae K.D. Hyde, Yin. Zhang, 

Suetrong & E.B.G. Jones 

Hyphopodia-like, anamorphic Trematosphaeria 

Fuckel, Jb. nassau. Ver. 

Naturk (C)* 

Pleosporales, genera incertae sedis 

Berkleasmium Zobel 1854 (H) 

Briansuttonia R.F. Castañeda, Minter & 

Saikawa 2004 (H) 

Cerebella Ces. 1851 (H) 

Cheiromoniliophora Tzean & J.L. Chen 

1990 (H) 

Diplococcium Grove 1885 (H) 

Ellisembia Subram. 1992 (H) 

Epicoccum Link 1815 (H) 

Fusculina Crous & Summerell 2006 (C) 

Mycocentrospora Deighton 1972 (H) 

Mycopappus Redhead & G.P. White 1985 

anamorphic Mycodidymella C.Z. Wei, 

Y. Harada & Katum. 1998 (H) 

Noosia Crous, R.G. Shivas & McTaggart 

2011 (H) 

Ochrocladosporium Crous & U. Braun 

2007 (H) 


Periconia Tode 1791(H) 

Pleurophomopsis Petr. 1924, anamorphic 

Astrosphaeriella Syd. & P. Syd. 1913 

(C) 

Polyschema H.P. Upadhyay 1966 (H) 

Pseudochaetosphaeronema Punith. 1979 

(C) 

Pseudopassalora Crous 2011 (H) 

Repetophragma Subram. 1992 (H)* 

?Scleroramularia Batzer & Crous 2011 

(H) 

Sclerostagonospora Höhn. 1917 (C) 

Sirodesmium De Not. 1849 (C) 

Spiroplana Voglmayr, M.J. Park & H.D. 

Shin 2011 (H) 

Sporidesmium Link 1809 (C) 

Versicolorisporium Sat. Hatak., Kaz. 

Tanaka & Y. Harada 2008 (C) 

Ybotromyces Rulamort 1986 (H) 

Dothideales genera incertae sedis 


Atramixtia Tsuneda, M.L. Davey & 

Currah 2011 (H) 

Celosporium Tsuneda & Davey 2010 (H) 

Diplococcium Grove 1885, anamorphic 

Otthia Nitschke ex Fuckel 1870 (H) 

Pleosporomycetidae, genera incertae sedis 

Acrogenospora M.B. Ellis 1971, anamorphic 

Farlowiella Sacc. 1891(H) 

Dothideomycetes, orders incertae sedis 

Acrospermales Minter, Peredo & A.T. Watson 

Acrospermaceae Fuckel 

Dactylaria Sacc. 1880, anamorphic Acrospermum 

Tode 1783 (H) 

Gonatophragmium Deighton 1969, anamorphic 

Acrospermum Tode 1790 (H) 

Botryosphaeriales C.L. Schoch, Crous & 

Shoemaker 

Botryosphaeriaceae Theiss. & H. Syd. 

Alysidiella Crous 2006 (H) 

Aplosporella Speg. 1880 (C) 

Arthrinium Kunze 1817, anamorphic Leptoguignardia 

E. Müll. (H) 

Bahusutrabeeja Subram. & Bhat 1977 (H) 

Barriopsis A.J.L. Phillips, A. Alves & 


Dichomera Cooke 1878 (C) 

Diplodia Fr. 1834 (C) 

Diplodia-like, anamorphic Neodeightonia 

C. Booth 1970 and Saccharata Denman 

& Crous 2004 (C) 

Dothichiza-like, anamorphic Leptoguignardia 

E. Müll. 1955 (C) 

Dothiorella Sacc. 1880, anamorphic Spencermartinsia 

A.J.L. Phillips, A. Alves 

& Crous 2008 (C) 

Fusicoccum Corda 1829, anamorphic 

Botryosphaeria Ces. & De Not. 1863 

(C) 

Fusicoccum-like, anamorphic Melanops 

Nitschke ex Fuckel 1870 and Saccharata 

Denman & Crous 2004 (C) 

Hormonema Lagerb. & Melin 1927, 

anamorphic Guignardia Viala & Ravaz 

1892 (H) 

Lasiodiplodia Ellis & Everh. 1896 (C) 

Leptodothiorella Höhn. 1918, anamorphic 

Guignardia Viala & Ravaz 1892 (C) 

Macrophomina Petr. 1923 (C) 

Microdiplodia Allesch. 1901 (C)

Neofusicoccum Crous, Slippers & A.J.L. 

Phillips 2006, with Botryosphaeria-like 

teleomorphs (C) 

Neoscytalidium Crous & Slippers 2006 

(H) 

Phyllosticta Pers. 1818, anamorphic Guignardia 

Viala & Ravaz 1892 (C) 

Pseudofusicoccum Mohali, Slippers & 

M.J. Wingf. 2006 (C) 

Sphaeropsis Sacc. 1880 (C) 

Thyrostroma Höhn. 1911, anamorphic 

Dothidotthia Höhn. 1918 (H) 

Botryosphaeriales, genera incertae sedis 

Camarosporium Schulzer 1870 (C) 

Dichomera Cooke 1878 (C) 

Hendersonula Speg. 1880 (C) 

Hysteriales Lindau 

Hysteriaceae Chevall. 

Aposphaeria-like, anamorphic Gloniopsis 

De Not. 1847 and Hysterobrevium E. 

Boehm & C.L. Schoch 2010 (C) 

Coniosporium Link 1809, anamorphic 

Hysterium Pers. 1797 (H) 

Plenodomus Preuss 1851, anamorphic 

Psiloglonium Höhn. 1918 (C) 

Septonema Corda 1837, anamorphic Oedohysterium 

E. Boehm & C.L. Schoch 

2009 (H) 

Sporidesmium Link 1809, anamorphic 

Psiloglonium Höhn. 1918 (H) 

Jahnulales Pang, Abdel-Wahab, El- 

Sharouney, E.B.G. Jones & Sivichai 

Aliquandostipitaceae Inderbitzin 

Brachiosphaera Nawawi 1976 (H) 

Mytilinidiales Boehm, C.L. Schoch & 

Spatafora 

Gloniaceae (Corda) Boehm, C.L. Schoch & 

Spatafora 

Cenococcum Moug. & Fr. 1829 (C) 

Cleistonium Speer 1986, anamorphic 

Glonium Mühl. 1813 (C) 

Mytilinidiaceae Kirschst. 

Camaroglobulus Speer 1986, anamorphic 

Mytilinidion Duby 1862 (C) 

Chalara (Corda) Rabenh. 1844, anamorphic 

Quasiconcha M.E. Barr & M. 

Blackw. 1981 (H)* 


?Monotosporella S. Hughes 1958, anamorphic 

Actidium Fr. 1815 (H) 

Papulospora Preuss 1851, anamorphic 

Lophium Fr. 1818 (H) 

Septonema Corda 1837, anamorphic 

Mytilinidion Duby 1862 (H) 

Taeniolella S. Hughes 1958, anamorphic 

Mytilinidion Duby 1861 (H) 

Jahnulales genera incertae sedis 

Xylomyces Goos, R.D. Brooks & Lamore 

1977 (H) 

Patellariales D. Hawksw. & O.E. Erikss. 

Patellariaceae Corda 

Aposphaeria-like, anamorphic Rhytidhysteron 

Speg. 1881 (C) 

Diplodia-like, anamorphic Rhytidhysteron 

Speg. 1881 (C) 

Venturiales Yin. Zhang, C.L. Schoch & K.D. 

Hyde 

Venturiaceae E. Müll. & Arx ex M.E. Barr 

Acroconidiellina (Berk. & Broome) M.B. 

Ellis 1971, anamorphic Acantharia 

Theiss. & Syd. (H)* 

Bactrodesmium Cooke 1883, anamorphic 

Phragmogibbera Samuels & Rogerson 

1990 (H) 

Dictyodochium Sivan. 1984, anamorphic 

Gibbera Fr. 1825 (H) 

?Didymochora Hohn. 1918, anamorphic 

Atopospora Petr. 1925 (C) 

Cladosporium-like, anamorphic Apiosporina 

Höhn. 1910 (H) 

Fusicladium Bonord. 1851, anamorphic 

Apiosporina Höhn. 1910, Caproventuria 

Braun 1998 and Venturia Sacc. 

1882 (H) 

Fusicladium-like, anamorphic Acantharia 

Theiss. & Syd 1918 and Protoventuria 

Berl. & Sacc. 1887 (H) 

Fusicladosporium Partr. & Morgan-Jones 

2003, anamorphic Venturia Sacc. 

1882 (H) 

Helicodendron Peyronel 1918 anamorphic 

Tyrannosorus Unter. & Malloch 1995 

(H) 

Pithosira Petr. 1949 (H) 

Spilocaea Fr. 1819, anamorphic Venturia 

Sacc. 1882 (C) 

Spilodochium Syd. 1927 anamorphic Pseu- 

167

168 

doparodiella F. Stevens 1927 (H) 

Stigmina-like, Sacc. 1980 anamorphic 

Aca-ntharia Theiss. & Syd 1918 and 

Gib-bera Fr. 1825 (H) 

Sympoventuriaceae Yin. Zhang, C.L. Schoch 

& K.D. Hyde 

Fusicladium-like (H) 

Sympodiella-like, anamorphic Sympoventuria 

Crous & Seifert (H)* 

Veronaeopsis Arzanlou & Crous 2007 

(H)* 

Dothideomycetes, families incertae sedis 

Ascoporiaceae Kutorga & D. Hawksw. 

Plectophomella-like, anamorphic Pseudosolidum 

Lloyd 1923 (H) 

Asterinaceae Hansf. 

?Acarella Syd. 1927, anamorphic Morenoina 

Theiss. 1913 (C) 

Asterostomella Speg. 1886, anamorphic 

Asterina Lév. 1845 (C) 

Asterostromina Bat. & A.F. Vital 1957, 

anamorphic Asterodothis Theiss. (C) 

Asterostomula Theiss. 1916, anamorphic 

Echidnodes Theiss. & Syd. 1918 (C) 

Bahusakala Subram. 1958 (H) 

?Clasterosporium Schwein. 1832, anamorphic 

Eupelte Syd. 1924 (H) 

Clasterosporium Schwein. 1832, anamorphic 

Asterina Lév. 1845, Asterodothis 

Theiss.1912, Yamamotoa Bat. 1960 and 

Lembosia Lév. 1845 (H) 

?Elachopeltis Syd. 1927, anamorphic Aphanopeltis 

Syd. 1927 (C) 

Eriothyrium Speg. 1888, anamorphic 

Aster-inema Bat. & Gayão 1953 (C)* 

Leprieurina G. Arnaud 1918, anamorphic 

Prillieuxina G. Arnaud 1918 (C) 

Mahanteshamomyces Hosag. & C.K. Biju 

2004 (C) 

?Peltasterella Bat. & H. Maia 1959, anamorphic 

Yamamotoa Bat. 1960 (C) 

Pirozynskia Subram. 1972, anamorphic 


Septoidium G. Arnaud 1921, anamorphic 


Septothyrella Höhn. 1911, anamorphic 

Uleothyrium Petr. 1929 (C) 

Shivamyces Hosag. 2004 (H) 


Sirothyriella Höhn. 1910, anamorphic 

Morenoina Thesis. 1913 (C) 

?Thyrinula Petr. & Syd. 1924, anamorphic 

Aulographina Arx & E. Müll. 1960 (C) 

Triposporium Corda 1837, anamorphic 

Batistinula Arx 1960 (H) 

Corynesporascaceae Sivan. 


Corynesporasca Sivan. (H) 

Dacampiaceae Körb. 

Cyclothyrium Petr. 1923, anamorphic Polycoccum 

Saut. ex Körb. 1865 (C) 

Lecythophora-like, anamorphic Munkovalsaria 

Aptroot 1995 (H) 

Englerulaceae Henn. 

Capnodiastrum Speg. 1886, anamorphic 

Englerula Henn. 1904 and Rhytidenglerula 

Höhn. 1918 (C) 

Digitosarcinella S. Hughes 1984, anamorphic 

Schiffnerula Höhn. 1909 (H) 

Mitteriella Syd. 1933, anamorphic Schiffnerula 

Höhn. 1909 (H) 

Questieriella G. Arnaud ex S. Hughes 

1983, anamorphic Schiffnerula Höhn. 

1909 (H) 

Sarcinella Sacc. 1880, anamorphic Schiffnerula 

Höhn. 1909 (H)* 

Eremomycetaceae Malloch & Cain 

Arthrographis G. Cochet ex Sigler & J.W. 

Carmich. 1976, anamorphic Eremomyces 

Malloch & Cain 1971and Pithoascina 

Valmaseda, A.T. Martínez & 

Barrasa 1987 (H) 

Trichosporiella Kamyschko 1960, anamorphic 

Rhexothecium Samson & 

Mouch. 1975 and Eremomyces Malloch 

& Cain 1971 (H) 

Euantennariaceae Hughes & Corlett 

Antennatula Fr. ex F. Strauss 1850, anamorphic 

Euantennaria Speg. and Strigopodia 

Bat. 1957 (H) 

Capnokyma S. Hughes 1975 (H) 

Capnophialophora S. Hughes 1966, anamorphic 

Strigopodia Bat. 1957 (H) 

Hormisciomyces Bat. & Nascim. 1957, 

anamorphic Euantennaria Speg. (H)

Hyphosoma Syd. 1924, anamorphic Strigopodia 

Bat. 1957 (H) 

Plokamidomyces Bat., C.A.A. Costa & 

Cif. 1957, anamorphic Trichopeltheca 

Bat., C.A.A. Costa & Cif. 1958 (H) 

Racodium Fr. 1829, anamorphic Strigopodia 

Bat. 1957 (H) 

Trichothallus F. Stevens 1925, anamorphic 

Trichopeltheca Bat., C.A.A. Costa & 

Cif. 1958 (H) 

Fenestellaceae M.E. Barr 


Fenestella Tul. & C. Tul. 1863 (C) 

Leptopeltidaceae Höhn. ex Trotter 

?Idriella P.E. Nelson & S. Wilh. 1956, 

anamorphic Dothiopeltis E. Mull. 1957 

(H) 

Leptothyrium-like, anamorphic Leptopeltis 

Höhn. 1917 (C) 

Meliolinaceae S. Hughes 

Briania D.R. Reynolds 1989, anamorphic 

Meliolina Syd. & P. Syd. 1914 (H) 

Micropeltidaceae Clem. & Shear 

Cyclopeltella Petr. 1953, anamorphic Cyclopeltis 

Petr. 1953 (C) 

Heliocephala V.G. Rao, K.A. Reddy & de 

Hoog 1984 (H)* 

Holubovaniella R.F. Castañeda 1985 (H)* 

Sirothyriella Höhn. 1910, anamorphic 

Stomiopeltis Theiss. 1914 (C) 

Sporidesmium-like, anamorphic Stomiopeltis 

Theiss. 1914 (as Akaropeltella 

M.L. Farr, 1972) (H) 

Pycnothyriaceous taxa associated with spe- 

cies of Stomiopeltis anamorphic Micropeltis 

Mont. 

Microthyriaceae Sacc. 

Asterostomula Theiss. 1916, anamorphic 

Asterinella Theiss. 1912 (C) 

Asteromella-like, anamorphic Asterinella 

Theiss. 1912 (C) 

Hansfordiella S. Hughes 1951, anamorphic 

Trichothyrium Speg. 1889 (H) 

Isthmospora F. Stevens 1918, anamorphic 

Trichothyrium Speg. 1889 (H) 

Leptothyrium Kunze 1823, anamorphic 


Microthyrium Desm. 1841 (C) 

Xenogliocladiopsis Crous & W.B. Kendr. 

1994, anamorphic Arnaudiella Petr. 

1927 (H) 

Zalerion R.T. Moore & Meyers 196, anamorphic 

Microthyrium Desm. 1841 (H) 

Parmulariaceae E. Müll. & Arx ex M.E. Barr 

Excipulariopsis P.M. Kirk & Spooner 

1982, anamorphic Kentingia Sivan. & 

W.H. Hsieh 1989 (H) 

?Melanoplaca Syd. & P. Syd. 1917, 

anamorphic Dothidasteroma Höhn. 

1909 (C) 

Placomelan Cif. 1962, anamorphic Dothidasteroma 

Höhn. 1909 (C) 

Parodiellaceae Theiss. & H. Syd. ex M.E. Barr 

?Ascochytopsis Henn. 1905, anamorphic 

Parodiella Speg. 1880 (C) 

Parodiopsidaceae Toro 

?Chuppia Deighton 1965, anamorphic 

Neoparodia Petr. & Cif. 1932 (H) 

Cicinnobella Henn. 1904, anamorphic 

Perisporiopsis Henn. 1904 (as Phaeodimeriella 

Speg. 1908) and Dimerium 

(Sacc. & P. Syd.) Sacc. & D. Sacc. 

1905 (C) 


Balladyna Racib. 1900 (H) 

Heterosporiopsis Petr. 1950, anamorphic 

‘Dimerium’ (Sacc. & P. Syd.) Sacc. & 

D. Sacc. 1905, Exosporinella Bender 

1932, possibly anamorphic Parodiellina 

Henn. ex G. Arnaud 1918 (H) 

?Ophiotrichum Kunze 1849 anamorphic 

Leptomeliola Höhn. 1919 (H) 

Sarcinella-like, anamorphic Neoparodia 

Petr. & Cif. 1932 (H) 

Septoidium G. Arnaud 1921, anamorphic 

Alina Racib. 1909, Perisporiopsis 

Henn. 1904, Parodiellina Henn. ex G. 

Arnaud 1918, Chevalieropsis G. 

Arnaud 1923, Ophioparodia Petr. & 

Cif. 1932 and Pilgeriella Henn. 1900 

(H) 

Shivamyces Hosag. 2004, anamorphic 

Alina Racib. 1909 (H) 

Tretospora M.B. Ellis 1976, anamorphic 

Balladyna Racib. and Balladynopsis 

169

170 

Theiss. & Syd. (H) 

Planistromellaceae M.E. Barr 

Aposphaeria-like, anamorphic Microcyclus 

Sacc., Syd. & P. Syd. 1904 (C) 

Fusicladium Bonord. 1851, anamorphic 

Microcyclus Sacc., Syd. & P. Syd. 1904 

(H) 

Hyphospora A.W. Ramaley 1996, anamorphic 

Comminutispora A.W. Ramaley 

1996 (H) 

Kellermania Ellis & Everh. 1885, anamorphic 

Planistroma A.W. Ramaley 

1991 (C) 

Lecanosticta Syd. 1922, anamorphic Eruptio 

M.E. Barr 1996 (C) 

Pazschkeella Syd. & P. Syd. 1901, anamorphic 

Microcyclus Sacc., Syd. & P. 

Syd. 1904 (C) 

Piptarthron Mont. ex Höhn. 1918, anamorphic 

Planistroma A.W. Ramaley 

1991 (C) 

Polystomellaceae Theiss. & H. Syd. 

Asteromella Pass. & Thüm. 1880 (as 

Stictochorella Höhn. 1917), anamorphic 

Dothidella Speg. 1880 (C) 

Lasmenia Speg. 1886, anamorphic Munkiella 

Speg. 1885 (C) 

Pseudoperisporiaceae Toro 

Chaetosticta Petr. & Syd. 1925, anamorphic 

Lasiostemma Theiss., Syd. & P. 

Syd. 1917 and ?Nematostoma Syd. & P. 

Syd. 1914 (C) 

Schizothyriaceae Höhn. ex Trotter, Sacc., D. 

Sacc. & Traverso 

Plenotrichaius Bat. & Valle 1961, anamorphic 

Mycerema Bat., J.L. Bezerra & 

Cavalc 1963 (C) 

Zygophiala E.W. Mason 1945, anamorphic 

Schizothyrium Desm. 1849 (H) 

Tubeufiaceae M.E. Barr 

?Annellospermosporella P.R. Johnst. 1999, 

anamorphic Malacaria Syd. 1930 (H) 

Aquaphila Goh, K.D. Hyde & W.H. Ho 

1998, anamorphic Tubeufia Penz. & 

Sacc. 1898 (H)* 


Araneomyces Höhn. 1909, anamorphic 

Paranectriella (P. Henn. & Sacc.) 

Höhn. (H) 

Chionomyces Deighton & Piroz. 1972, 

anamorphic Melioliphila Speg. 1924 

(H) 


Chaetosphaerulina I. Hino 1938 (H) 

Eriomycopsis Speg. 1910, anamorphic 

Melioliphila Speg. 1924 and Byssocallis 

Syd. 1927 (H) 

Guelichia Speg. 1886, anamorphic Puttemansia 

Henn. 1902 (H) 

Helicoma Corda 1837, anamorphic Tubeufia 

Penz. & Sacc. 1898, Thaxteriella 

Petr. 1924 and Chlamydotubeufia Boonmee 

& K.D. Hyde 2011 (H)* 

Helicoön Morgan 1892 (H)* 

Helicomyces Link 1809, anamorphic 

Acanthostigma De Not. (1863) (H)* 

Helicosporium Nees 1816, anamorphic 

Acanthostigma De Not. (1863) and 

Tubeufia Penz. & Sacc. 1898 (H) 

Kamalomyces R.K. Verma, N. Sharma & 

Soni 2008 (C) 

Mirandina G. Arnaud ex Matsush. 1975, 

anamorphic Taphrophila Scheuer 1988 

(H)* 

Monodictys S. Hughes 1958, anamorphic 


Monodictys-like, anamorphic Tubeufia 

Penz. & Sacc. 1898 (H) 

Moorella P. Rag. Rao & D. Rao 1964, 

anamorphic Chaetosphaerulina I. Hino 

1938 (H) 

Pendulispora M.B. Ellis 1961, anamorphic 


Peziotrichum (Sacc.) Lindau 1900, anamorphic 

Podonectria Petch 1921 (H) 

Spermatoloncha Speg. 1908 (H) 

Tetracrium Henn. 1902, anamorphic Puttemansia 

Henn. 1902 and Podonectria 

Petch 1921 (H) 

Titaea Sacc. 1876, anamorphic Paranectriella 

(Henn. ex Sacc. & D. Sacc.) 

Höhn. 1910 and Puttemansia Henn. 

1902 (H) 

Xenosporium Penz. & Sacc. 1901, anamorphic 

Acanthostigmella Höhn. 1905 and 

Chaetosphaerulina I. Hino 1938 (as

Thaxteriellopsis Sivan., Panwar & S.J. 

Kaur 1977) (H) 

Vizellaceae H.J. Swart 

Chrysogloeum Petr. 1959, anamorphic 

Blasdalea Sacc. & P. Syd. 1902 (C) 

Manginula G. Arnaud 1918, anamorphic 

Vizella Sacc. 1883 (C) 

Dothideomycetes, genera incertae sedis 

Acrocalymma Alcorn & J.A.G. Irwin 

(1987), anamorphic Massarina, however 

Massarina has been shown to be 

polyphyletic and its placement is therefore 

presently unclear (C) 

Ampullifera Deighton 1960, anamorphic 

Teratoschaeta Bat. & O.M. Fonseca 

(H) 

Ascochyta-like, anamorphic Gilletiella 

Sacc. & P. Syd. 1899 (C) 

Asteromella Pass. & Thüm. 1880 (C) 

Bactrodesmium Cooke 1883, anamorphic 

Stuartella Fabre 1879 (H) 

Brachyconidiella R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Camarosporula Petr. 1954, anamorphic 

Anthracostroma Petr. 1954 (C) 

Chaetasbolisia Speg. 1918, anamorphic 

Pseudomorfea Punith. 1981 (C) 

Cladoriella Crous 2006 (H) 

Coniosporium Link 1809 (H) 

Coronospora M.B. Ellis 1971, anamorphic 

Ascocoronospora Matsush. 2003 (H) 

Cryomyces Selbmann, de Hoog, Mazzaglia, 

Friedmann & Onofri 2005 (H) 

?Cyclothyrium Petr. 1923, anamorphic 

Thyridaria Sacc. 1875 (C) 

Dendryphiopsis S. Hughes 1953, anamorphic 

Kirschsteiniothelia D. Hawksw. 

1985 (H) 

Diederichia D. Hawksw. 2003 (C) 

Dilophospora Desm. 1840, anamorphic 

Lidophia J. Walker & B. Sutton 1974 

(C) 

Diplodia-like, anamorphic Otthia Nitschke 

ex Fuckel 1870 (C) 

Disculina Höhn. 1916 (C) 

Funbolia Crous & Seifert 2011 (H) 

Hansfordiellopsis Deighton 1960, anamorphic 

Koordersiella Höhn. 1909 (H) 


Helicodendron Peyronel 1918, anamorphic 

Tyrannosorus Unter. & Malloch 1995 

(H) 

Hiospira R.T. Moore 1962, anamorphic 

Brooksia Hansf. 1956 (H) 

Iledon Samuels & J.D. Rogers 1986, anamorphic 

Botryohypoxylon Samuels & 

J.D. Rogers 1986 (C) 

Megaloseptoria Naumov 1925 (C) 


anamorphic Chaetonectrioides Matsush. 

1996 (H) 

Monodictys S. Hughes 1958 (H) 

Peltaster Syd. & P. Syd. 1917 (C) 

Phaeothecoidiella Batzer & Crous 2010 

(H) 

Phanerococculus Cif. 1954, anamorphic 

Koordersiella Höhn. 1909 (C) 

Phialophora-like, anamorphic Tirisporella 

E.B.G. Jones, K.D. Hyde & Alias 1996 

(H) 

Phoma-like, anamorphic Tremateia Kohlm., 

Volkm.-Kohlm. & O.E. Erikss. 

1995 (C) 

Placosphaeria (De Not.) Sacc. 1880 (C) 

Plectopycnis Bat. & A.F. Vital 1959 (C) 


Gibberidea Fuckel 1870 (C) 

Pseudorobillarda M. Morelet 1968 (C) 

Pycnopleiospora C.Z. Wei, Y. Harada & 

Katum. 1997, anamorphic Pseudodidymella 

C.Z. Wei, Y. Harada & Katum. 

1997 (H) 

Rhizopycnis D.F. Farr 1998 (C) 

Scleroconidioma Tsuneda, Currah & Thormann 

2000 (H) 

Septoriella Oudem. 1889 (C) 

Troposporella P. Karst. 1892 (H) 

Class Eurotiomycetes Tehler ex O.E. Eriksson 

& K. Winka 

Subclass Chaetothyriomycetidae Doweld 

Chaetothyriales M.E. Barr 

Chaetothyriaceae Hansf. ex M.E. Barr 

?Ciferrioxyphium Bat. & H. Maia 1963, 

anamorphic Aithaloderma P. Syd. (H)* 

Cyphellophora G.A. de Vries 1962 (H) 

Leptoxyphium Speg. 1918, anamorphic Aithaloderma 

P. Syd. 1913 (H)* 

Merismella Syd. 1927, anamorphic Chaet- 

171

172 

othyrium Speg. 1888 (C) 

Stanhughesia Constant. 1989, anamorphic 

Ceramothyrium Bat. & H. Maia 1956 

(H) 

Tripospermum Speg. 1918, anamorphic 

Phragmocapnias Theiss. & Syd. 1918, 

Trichomerium Speg. 1918 and Triposporiopsis 

W. Yamam. 1955 (H)* 

Vonarxia Bat. 1960 (H) 

Herpotrichiellaceae Munk 

Brycekendrickomyces Crous & M.J. 

Wingf. 2009 (H) 

Cladophialophora Borelli 1980, anamorphic 

Capronia Sacc. 1883 (H) 

Exophiala J.W. Carmich. 1966, anamorphic 


Fonsecaea Negroni 1936, anamorphic 


Metulocladosporiella Crous, Schroers, J.Z. 

Groenew., U. Braun & K. Schub. 2006 

(H) 

Nadsoniella Issatsch. 1914 (H) 

Phaeococcomyces de Hoog 1979 (H) 

Phaeoramularia Munt.-Cvetk. 1960, anamorphic 

Capronia Sacc.1883 (H) 

Phialophora Medlar 1915 (H) 

Racodium Pers. 1794, anamorphic Capronia 

Sacc. 1883 (H) 

Rhinocladiella Nannf. 1934, anamorphic 


Sorocybe Fr. 1849 (H) 

Thysanorea Arzanlou, W. Gams & Crous 

2007 (H) 

Veronaea Cif. & Montemart. 1957 (H) 

Chaetothyriales M.E. Barr, genera incertae 

sedis 

Coniosporium Link 1809, the type species 

is considered as Sirodesmium De Not. 

1849 (H) 

Phaeoannellomyces McGinnis & Schell 

1985 (H)(= Hortaea) 

Staninwardia B. Sutton 1971 (C) 

Strelitziana Arzanlou & Crous 2006 (H) 

Pyrenulales, genera incertae sedis 

Lyromma Bat. & H. Maia 1965 (C) 

Chaetothyriomycetidae, families incertae sedis 

Rhynchostomataceae Winka & O. E. Erikss. 


Arthropycnis Constant. 1992, anamorphic 

Rhynchostoma P. Karst. (C) 

Strigulaceae Zahlbr. 

Discosiella Syd. & P. Syd. 1912, anamorphic 

Strigula Fr. 1823 (C) 

Chaetothyriomycetidae, genera incertae sedis 

Peyronelia Cif. & Gonz. Frag. 1927, anamorphic 

Glyphium Nitschke ex F. 

Lehm. 1886 (H) 

Phaeomoniella Crous & W. Gams 2000 

(H) 

Rhabdospora-like, anamorphic Dolabra C. 

Booth & W.P. Ting 1964 (C) 

Taeniolella S. Hughes 1958, anamorphic 

Glyphium Nitschke ex F. Lehm. 1886 

(H) 

Subclass Eurotiomycetidae Geiser & Lutzoni 

Coryneliales Seaver & Chardon 

Coryneliaceae Sacc. ex Berl. & Voglino 

Exophiala J.W. Carmich. 1966, anamorphic 

Bicornispora Checa, Barrasa, 

M.N. Blanco & A.T. Martínez 1996 (H) 

Eurotiales G.W. Martin ex Benny & Kimbr. 

Monascaceae J. Schröt. 

Basipetospora G.T. Cole & W.B. Kendr. 

1968, anamorphic Monascus Tiegh. 

1884 (H) 

Fraseriella Cif. & A.M. Corte 1957, anamorphic 

Xeromyces Fraser (H) 

Trichocomaceae E. Fisch. 

Aspergillus P. Micheli ex Link 1809, 

anamorphic Dichlaena Durieu & Mont. 

1849, Emericella Berk. & Broome 

1857, Eurotium Link 1809, Fennellia 

B.J. Wiley & E.G. Simmons 1973, 

Hemicarpenteles A.K. Sarbhoy & 

Elphick 1968, Neopetromyces Frisvad 

& Samson 2000, Neosartorya Malloch 

& Cain 1973, Petromyces Malloch & 

Cain 1973 and Warcupiella Subram. 

1972 (H)* 

Geosmithia J.Pitt 1979, anamorphic, Chromocleista 

Yaguchi & Udagawa 1993 

and Talaromyces C.R. Benj. 1955 (H) 

Isaria Pers. 1794, anamorphic Byssochlamys 

Westling (H)

Merimbla Pitt 1979, anamorphic Talaromyces 

C.R. Benj. 1955 (H) 

Paecilomyces Bainier 1907, anamorphic 

Byssochlamys Westling 1909 and Talaromyces 

C.R. Benj. 1955 (H) 

Paecilomyces-like, anamorphic Coonemeria 

Mouch. 1997 (H) 

Penicillium Link 1809, anamorphic Eupenicillium 

F. Ludw. 1892, Talaromyces 

C.R. Benj. 1955, Dichlaena Durieu & 

Mont. 1849 and Trichocoma Jungh. 

1838 (H) 

Phialosimplex Sigler, Deanna A. Sutton, 

Gibas, Summerb. & Iwen 2010 (H) 

Polypaecilum G. Sm. 1961, anamorphic 

Dactylomyces Sopp 1912, Dichotomomyces 

Saito ex D.B.Scott 1970 and 

Thermoascus Miehe 1907 (H) 

Pseudocordyceps Hauman 1936, anamorphic 

Penicilliopsis Solms 1887 (H) 

Sagenomella W. Gams 1978, anamorphic 

Sagenoma Stolk & G.F. Orr 1974 and 

Talaromyces C.R. Benj. 1955 (H) 

Sarophorum Syd. & P. Syd. 1916, anamorphic 


Stilbodendron Syd. & P. Syd. 1916, anamorphic 


Stilbothamnium Henn. 1896 (H) 

Thermomyces Tsikl. 1899 (H) 

Thysanophora W.B. Kendr. 1961 (H) 

Torulomyces Delitsch 1943, anamorphic 

Eupenicillium F. Ludw. 1892 (H) 

Eurotiomycetidae genera incertae sedis 

Penicillium Link 1809, anamorphic Paratalaromyces 


Onygenales Cif. ex Benny & Kimbr. 

Ajellomycetaceae Untereiner, J.A. Scott & 

Sigler 

Blastomyces Gilchrist & W.R. Stokes 

1898, anamorphic Ajellomyces McDonough 

& A.L. Lewis 1968 (H) 

Chrysosporium Corda 1833, anamorphic 

Ajellomyces McDonough & A.L. Lewis 

1968 (H) 

Emmonsia Cif. & Montemart. 1959, anamorphic 

Ajellomyces McDonough & 

A.L. Lewis 1968 (H) 

Histoplasma Darling 1906, anamorphic 

Ajellomyces McDonough & A.L. Lewis 

1968 (H) 


Lacazia Taborda, V.A. Taborda & Mc- 

Ginnis 1999 (H) 

Paracoccidioides F.P. Almeida 1930 (H) 

Arachnomycetaceae Gibas, Sigler & Currah 

Onychocola Sigler 1990, anamorphic 

Arachnomyces Massee & E.S. Salmon 

1902 (H) 

Gymnoascaceae Baran. 

Chrysosporium-like, anamorphic Orromyces 

B. Sur & G.R. Ghosh 1987 (H) 

Malbranchea Sacc. 1882, anamorphic 

Byssoonygena Guarro, Punsola & Cano 

1987 and Gymnoascus Baran. 1872 (H) 

Oncocladium Wallr. 1833, anamorphic 

Gymnoascus Baran. 1872 (H) 

Onygenaceae Berk. 

?Arthrographis G. Cochet ex Sigler & 

J.W. Carmich. 1976, anamorphic Arachnotheca 

Arx 1971 (H) 

Chrysosporium Corda 1833 anamorphic 

Amauroascus J. Schröt., Aphanoascus 

Zukal 1890, Apinisia La Touche 1968, 

Arachnotheca Arx 1971, Nannizziopsis 

Currah 1985, ?Polytolypa J.A. Scott & 

Malloch 1993, Pectinotrichum Varsavsky 

& G.F. Orr 1971, ?Renispora Sigler 

& J.W. Carmich. 1979 (H) 

Coccidioides G.W. Stiles 1896 (H) 

Malbranchea Sacc. 1882, anamorphic Auxarthron 

G.F. Orr & Kuehn 1963, 

Byssoonygena Guarro, Punsola & Cano 

1987 and Uncinocarpus Sigler & G.F. 

Orr 1976 (H) 

Myriodontium Samson & Polon. 1978, 

anamorphic Neoarachnotheca Ulfig, 

Cano & Guarro 1997 (H) 

Sporendonema Desm. 1827, anamorphic 

Onygena Pers. 1800 (H) 

Arthrodermataceae Currah 


Shanorella R.K. Benj. 1956, Arthroderma 

Curr. 1860 and Ctenomyces 

Eidam 1880 (H) 


Epidermophyton Sabour. 1907 (H) 

Microsporum Gruby 1843, anamorphic 

Nannizzia Stockdale 1961 and Arthroderma 

Curr. 1860 (H) 

173

174 

Myceliophthora Costantin 1892, anamorphic 

Arthroderma Curr. 1860 

Trichophyton Malmsten 1848, anamorphic 

Arthroderma Curr. 1860 (H) 

Ascosphaeraceae L.S. Olive & Spiltoir 

Chrysosporium-like, anamorphic Ascosphaera 

L.S. Olive & Spiltoir 1955 and 

Bettsia Skou 1972 (H) 

Onygenales, genera incertae sedis 

Arthropsis Sigler, M.T. Dunn & J.W. 


Lacazia Taborda, V.A. Taborda & McGinnis 

1999 (H) 

Subclass Mycocaliciomycetidae Tibell 

Mycocaliciales Tibell & Wedin 

Mycocaliciaceae A.F.W. Schmid 

Asterophoma D. Hawksw. 1981, anamorphic 

Chaenothecopsis Vain. 1927 (C) 

Catenomycopsis Tibell & Constant. 1991, 

anamorphic Chaenothecopsis Vain. 

1927 (H) 

Phialophora-like, anamorphic Chaenothecopsis 

Vain. 1927 and Mycocalicium 

Vain. 1890 (H) 

Class Laboulbeniomycetes Engler 

Laboulbeniales Engler 

Coreomycetopsis Thaxt. 1920 (H) 

Muiaria Thaxt. 1914 (H) 

Tetrameronycha Speg. ex W. Rossi & M. 

Blackw. 1990 (H) 

Pyxidiophorales P.F. Cannon 

Pyxidiophoraceae Arnold 

Chalara (Corda) Rabenh. 1844 anamorphic 

Pyxidiophora Bref. & Tavel 

1891 (H)* 

Gliocephalis Matr. 1899 (H) 

Pleurocatena G. Arnaud ex Aramb., 

Gamundí, W. Gams & G.R.W. Arnold 

2007, anamorphic Pyxidiophora Bref. 

& Tavel 1891 (H) 

Thaxteriola Speg. 1918, anamorphic Pyxidiophora 

Bref. & Tavel 1891 (H) 

Class Lecanoromycetes O.E. Erikss. & Winka 

Subclass Ostropomycetidae Reeb, Lutzoni & 

Cl. Roux 


Agyriales Clem. & Shear 

Trapeliaceae M. Choisy ex Hertel 

Epithyrium (Sacc.) Trotter 1931, anamorphic 

Sarea Fr. 1825 (C) 

Pycnidiella Höhn. 1915, anamorphic Sarea 

Fr. 1825 (C) 

Ostropales Nannf. 

Odontotremataceae D. Hawksw. & Sherwood 

Pycnidial anamorph associated with 

Claviradulomyces P.R. Johnst., D.C. 

Park, H.C. Evans, R.W. Barreto & D.J. 

Soares 2010 

Stictidaceae Fr. 

?Coleophoma-like, anamorphic Stictophacidium 

Rehm 1888 (C) 

?Ebollia Minter & Caine 1980, anamorphic 

Stictophacidium Rehm 1888 (C) 

Phacidiella-like, anamorphic Acarosporina 

Sherwood 1977 (C) 

Stictospora Cif. 1957, anamorphic Stictis 

Pers. 1800 (C) 

Ostropales, genera incertae sedis 

Glomerobolus Kohlm. & Volkm.-Kohlm. 

1996 (H) 

Subclass Lecanoromycetidae P.M. Kirk, P.F. 

Cannon, J.C. David & Stalpers ex Miadl., 

Lutzoni & Lumbsch 

Lecanorales Nannf. 

Pilocarpaceae Zahlbr. 

Pyriomyces Bat. & H. Maia 1965, 

anamorphic Byssoloma Trevis. 1853 

(C) 

Szczawinskia A. Funk 1984 (C) 

Stereocaulaceae Chevall. 

Lepraria Ach. 1803 (C) 

Lecanorales, genera incertae sedis 

Leprocaulon Nyl. 1878 (C) 

Vouauxiomyces Dyko & D. Hawks. 1979, 

anamorphic Abrothallus De Not. 1845 

(C) 

Class Leotiomycetes O.E. Erikss. & Winka 

Cyttariales Luttr. ex Gamundı´ 

Cyttariaceae Speg. 

Cytariella Palm 1932, anamorphic Cytta-

ia Berk. 1842 (C) 

Erysiphales Gwynne-Vaughan 

Erysiphaceae Tul. & C. Tul. 

Oidiopsis Scalia 1902, anamorphic Leveillula 

G. Arnaud 1921 (H) 

Oidium Link 1824, anamorphic Arthrocladiella 

Vassilkov 1960, Blumeria 

Golovin ex Speer 1975, Brasiliomyces 

Viégas 1944, Cystotheca Berk. & M.A. 

Curtis 1860, Erysiphe R. Hedw. ex DC. 

1805, Golovinomyces (U. Braun) V.P. 

Gelyuta 1988, Medusosphaera Golovin 

& Gamalizk. 1962, Microsphaera Lév. 

1851, U. Braun 1999, Podosphaera 

Kunze 1823, Sawadaea Miyabe 1914, 

Sphaerotheca Lév. 1851, Uncinula Lév. 

1851 and Uncinuliella R.Y. Zheng & 

G.Q. Chen 1979 (H) 

Ovulariopsis Pat. & Har. 1900, anamorphic 

Phyllactinia Lév. 1851 (H) 

Streptopodium R.Y. Zheng & G.Q. Chen 

1978, anamorphic Pleochaeta Sacc. & 

Speg. 1881 and Phyllactinia Lév. 1851 

(H) 

Helotiales Nannf. 

Bulgariaceae Fr. 

Crinula Fr. 1821, anamorphic Holwaya 

Sacc. 1889 (H) 

Endomelaconium Petr. 1940, anamorphic 

Bulgaria Fr. 1822 (C) 

Dermateaceae Fr. 


Mollisia (Fr.) P. Karst. 1871 (H) 

Cadophora Lagerb. & Melin 1927, anamorphic 

Mollisia (Fr.) P. Karst. 1871, 

Pyrenopeziza Fuckel 1870 and Pseudopezicula 

Korf 1975 (H) 

Cadophora-like, anamorphic Phaeomollisia 

T.N. Sieber & Grünig 2009 (H) 

Casaresia Gonz. Frag. 1920, anamorphic 


Chondropodium-like, anamorphic Waltonia 

Saho 1970 (C) 

Corniculariella P. Karst. 1884, anamorphic 

Dermea Fr. 1825 and Durandiella 

Seaver 1932 (C) 

Cryptocline Petr. 1924, anamorphic 

Trochila Fr. 1849 (C) 

Cryptosporiopsis Bubák & Kabát 1912, 

anamorphic Neofabraea H.S. Jacks. 


1913, Pezicula Tul. & C. Tul. 1865 and 

?Ocellaria (Tul. & C. Tul.) P. Karst. 

1871 (C) 

Cryptosympodula Verkley 1999, anamorphic 

Scleropezicula Verkley 1999 (C) 

Cylindrocolla Bonord. 1851, anamorphic 

Calloria Fr. 1836 (H) 

Cylindrosporium Grev. 1822, anamorphic 

Pyrenopeziza Fuckel 1870 (C) 

Cystodendron Bubák 1914, anamorphic 

Mollisia (Fr.) P. Karst. 1871 Belonopsis 

(Sacc.) Rehm 1891 and Niptera Fr. 

1849 (H) 

Diplosporonema Höhn. 1917, anamorphic 


Discogloeum Petr. 1923, anamorphic 

Spilopodia Boud. 1885 (C) 

Entomosporium Lév. 1856, anamorphic 

Diplocarpon F.A. Wolf 1912 (C) 

Eriosporella Höhn. 1916, possibly anamorphic 

Laetinaevia Nannf. 1932 (C) 

Foveostroma DiCosmo 1978, anamorphic 

Dermea Fr. 1825 (C) 

Gelatinosporium Peck 1873, anamorphic 

Dermea Fr. 1825 and Durandiella 

Seaver 1932 (C) 

Glarea Bills & Paláez 1999 (H) 

Gloeosporidiella Petr. 1921, anamorphic 

Drepanopeziza (Kleb.) Höhn. 1917 and 

Pseudopeziza Fuckel 1870 (C) 

Hainesia Ellis & Sacc. 1884, anamorphic 

Discohainesia Nannf. 1932 (C) 



Laetinaevia Nannf. 1932, anamorphic 

Naeviopsis B. Hein 1976 (C) 

Marssonina Magnus 1906, anamorphic 

Diplocarpon F.A. Wolf 1912 & 

Drepanopeziza (Kleb.) Höhn. 1917 (C) 

Melanodiscus Höhn. 1918, anamorphic 

Spilopodia Boud. 1885 (H) 

Microgloeum Petr. 1922, anamorphic 

Blumeriella Arx 1961 (C) 

Monostichella Höhn. 1916, anamorphic 

Drepanopeziza (Kleb.) Höhn. 1917 (C) 

Myrioconium Syd. & P. Syd. 1912, anamorphic 

Dibeloniella Nannf. 1932 (H) 

Phacidiella P. Karst. 1884, anamorphic 


Phialocephala W.B. Kendr. 1961 

anamorphic Phaeomollisia T.N. Sieber 

& Grünig 2009 (H) 

175

176 

Phialocephala-like, anamorphic Phaeomollisia 

T.N. Sieber & Grünig 2009 

(H) 

Phialophora-like, anamorphic Dibeloniella 

Nannf. 1932 (H) 

Phloeosporella Höhn. 1924, anamorphic 

Blumeriella Arx 1961 (C) 

Phlyctema Desm. 1847, anamorphic Neofabraea 

H.S. Jacks. 1913 (C) 

Pilidium Kunze 1823, anamorphic Discohainesia 

Nannf. 1932 (C) 

Rhynchosporium Heinsen ex A.B. Frank 

1897 (H)* 

?Sporonema Desm. 1847, anamorphic 

Leptotrochila P. Karst. 1871 (C) 


Laetinaevia Nannf. 1932 (H) 

Trimmatostroma Corda 1837 (H) 

Xenochalara M.J. Wingf. & Crous 2000 

(H) 

Helotiaceae Rehm 

Alatospora Ingold 1942 (H) 


Hymenoscyphus Gray 1821 (H) 

Articulospora Ingold 1942, anamorphic 

Hymenoscyphus Gray 1821(H) 

Ascoconidium Seaver 1942, anamorphic 

Sageria A. Funk 1975 (H) 

Aureohyphozyma Hosoya & Y. Otani 1995 

anamorph of Gelatinipulvinella Hosoya 

& Y. Otani (Hosoay Y Otani 1995) (H) 

Bloxamia Berk. & Broome 1854, anamorphic 

Bisporella Sacc. 1884 (H) 

Bothrodiscus Shear 1907, anamorphic 

Ascocalyx Naumov 1926 (C) 

Brunchorstia Erikss. 1891, anamorphic 

Gremmeniella M. Morelet 1969 (C) 

?Bryophytomyces Cif. 1953, anamorphic 


Cercosporella Sacc. 1880, anamorphic 

Heterosphaeria Grev. 1824 (H) 

Coryne Nees 1816, anamorphic Ascocoryne 

J.W. Groves & D.E. Wilson 

1967 (H) 

Cystodendron Bubák 1914, anamorphic 

Bisporella Sacc. 1884 (H) 

?Cystotricha Berk. & Broome 1850, anamorphic 

Xylogramma Wallr. 1833 (C) 

Deltosperma W.Y. Zhuang 1988, anamorphic 

Unguiculariopsis Rehm 1909 


(C) 

Dendrostilbella Höhn. 1905, anamorphic 

Claussenomyces Kirschst. 1923 (H) 

Dichaenopsella Petr. 1952, type is 

anamorphic Godroniopsis Diehl & E.K. 

Cash 1929 (C) 

Digitosporium Gremmen 1953, anamorphic 

Crumenulopsis J.W. Groves 

1969 (C) 

Dimorphospora Tubaki 1958, anamorphic 


Dothiorina Höhn. 1911, anamorphic Chlorociboria 

Seaver ex Ramamurthi, Korf 

& L.R. Batra 1958 (C) 

Endomelaconium Petr. 1940, anamorphic 

Austrocenangium Gamundí 1997 (C) 

Eriomycopsis Speg. 1910, anamorphic 

Calloriopsis Syd. & P. Syd. 1917 (H) 

Eustilbum Rabenh. 1864, anamorphic Bisporella 

Sacc. 1884 (H) 

Fuckelia Bonord. 1864, anamorphic Godronia 

Moug. & Lév. 1846 (C) 

Geniculospora Sv. Nilsson ex Marvanová 

& Sv. Nilsson 1971, anamorphic Hymenoscyphus 

Gray 1821 (H) 

Gliomastix Guég. 1905 anamorphic 

Ascoclavulina Y. Otani 1974 (H)* 

Heteropatella Fuckel 1874, anamorphic 

Heterosphaeria Grev. 1824 (C) 

Hormonema-like, anamorphic Pragmopora 

A. Massal 1855 (H) 

Idriella P.E. Nelson & S. Wilh. 1956, 

anamorphic Hymenoscyphus Gray 1821 

(H) 

Linodochium Höhn. 1909, anamorphic 

Pseudohelotium Fuckel 1870 (H) 

Monodictys S. Hughes 1958 anamorphic 


Periperidium Darker 1963, anamorphic 

Micraspis Darker 1963 (C) 

Phialophora-like, anamorphic Pseudopezicula 

Korf 1975 (H) 

Pragmopycnis B. Sutton & A. Funk 1975, 

anamorphic Pragmopora A. Massal 

1855 (C) 

?Pseudaegerita J.L. Crane & Schokn. 

1981, anamorphic Claussenomyces Kirschst. 

1923 (H) 

Pseudospiropes M.B. Ellis 1971, anamorphic 

Strossmayeria Schulzer 1881 

(H)

Rhinocladiella-like, anamorphic Bioscypha 

Syd. 1927 (H) 

Rhizothyrium Naumov 1915, anamorphic 

Rhizocalyx Petr. 1928 (C) 

Scytalidium Pesante 1957 anamorphic 


Sirodothis Clem. 1909, anamorphic Tympanis 

Tode 1790 (C) 

Sporonema Desm. 1847, anamorphic Godronia 

Moug. & Lév. 1846 (C) 

Symphyosirella Seifert 2009, anamorphic 

Symphyosirinia E.A. Ellis 1956 (H) 

Titaeospora Bubák 1916, anamorphic 

Stamnaria Fuckel 1870 (C) 

Topospora Fr. 1836, anamorphic Godronia 

Moug. & Lév. 1846 (C) 

Tricladium Ingold 1942, anamorphic 

Cudoniella Sacc. 1889 and Hymenoscyphus 

Gray 1821 (H) 

Varicosporium W. Kegel 1906, anamorphic 


Hemiphacidiaceae Korf 

Meria Vuill. 1896, anamorphic Rhabdocline 

Syd. 1922 (H) 

Rhabdogloeum Syd. 1922, anamorphic 

Rhabdocline Syd. 1922 (C) 

Rhabdogloeopsis Petr. 1925, anamorphic 

Sarcotrochila Höhn. 1917 (C) 

Hyaloscyphaceae Nannf. 

Acleistia Bayl. Ell. (1917) anamorphic 

Calycina Nees ex Gray 1821 (C) 

Brefeldochium Verkley 2005, anamorphic 

Polydesmia Boud. 1885 (H) 

Chaetochalara B. Sutton & Piroz. 1965 

(H) 


Calycellina Höhn., Phaeoscypha 

Spooner 1984 and Tapesina 

Lambotte 1887 (H)* 

Cheiromycella Höhn. 1910, anamorphic 

Hyaloscypha Boud. 1885 (H) 

Clathrosphaerina Beverw. 1951, anamorphic 

Hyaloscypha Boud. 1885 (H) 

Haplographium Berk. & Broome 1859, 

anamorphic Dematioscypha Svrček 

1977 (H) 

Mycoarthris Marvanová & P.J. Fisher 

2002 (H) 


Naemospora Roth ex Kuntze 1898, 

possibly anamorphic Lachnellula P. 

Karst. 1884 (C) 

Phialophora-like, anamorphic Cistella 

Quél. and Hyalopeziza Fuckel 1870 (H) 

Pseudaegerita J.L. Crane & Schokn. 1981, 

anamorphic Hyaloscypha Boud. 1885 

(H) 

Septonema-like, anamorphic Ciliolarina 

Svrcek 1977 (H) 

Tricladium Ingold 1942, anamorphic 

Hydrocina Scheuer 1991 (H) 

Leotiaceae Corda 


Pezoloma Clem. 1909 (H) 

Aureohyphozyma Hosoya & Y. Otani 

1995, anamorphic Gelatinipulvinella 

Hosoya & Y. Otani 1995 (H) 

Halenospora E.B.G. Jones 2009 (H) 

Myrioconium-like, anamorphic Neobulgaria 

Petr. 1921 (H) 

Phialophora-like, anamorphic Neobulgaria 

Petr. 1921 (H) 

Loramycetaceae Dennis ex Digby & Goos 


Loramyces W. Weston 1929 (H) 

Phacidiaceae Fr. 

Allantophomopsis Petr. 1925, anamorphic 

Phacidium Fr. 1815 (C) 

Apostrasseria Nag Raj 1983, anamorphic 

Lophophacidium Lagerb. 1949 and 


Blennoria Moug. & Fr. 1825, anamorphic 


Ceuthospora Grev. 1826, anamorphic 


?Coma Nag Raj & W.B. Kendr. 1972, 

anamorphic Ascocoma H.J. Swart (C) 

Rutstroemiaceae Holst-Jensen, L.M. Kohn & 

T. Schumacher 


Lambertella Höhn. 1918 (H) 

Myrioconium Syd. & P. Syd. 1912, ?anamorphic 

Lanzia Sacc. 1884, Rutstroemia 

P. Karst. 1871, Scleromitrula 

177

178 

S.Imai 1941, Lambertella Höhn. 1918 

and Poculum Velen. 1934 (H) 

Phialophora-like, anamorphic Rutstroemia 

P. Karst. 1871 (H) 

Sclerotiniaceae Whetzel ex Whetzel 

Acarosporium Bubák & Vleugel ex Bubák 

1911, anamorphic Pycnopeziza W.L. 

White & Whetzel 1938 (C) 

Amerosporium Speg. 1882, anamorphic 

Zoellneria Velen. 1934 (C) 

Amphobotrys Hennebert 1973, anamorphic 

Botryotinia Whetzel 1945 (H) 

Botrytis P. Micheli ex Pers. 1794, 

anamorphic Botryotinia Whetzel 1945 

(H) 

Cristulariella Höhn. 1916, anamorphic 

Nervostroma Narumi & Y. Harada 

2006 (H) 

Haradamyces Masuya, Kusunoki, Kosaka 

& Aikawa 2009 (H) 

Hinomyces Narumi & Y. Harada 2006, 

anamorphic Grovesinia M.N. Cline, 

J.L. Crane & S.D. Cline 1983 (H) 

Monilia Bonord. 1851, anamorphic Monilinia 

Honey 1928 and Phaeosclerotinia 

Hori 1916 (H) 

Mycopappus Redhead & G.P. White 1985, 

anamorphic Redheadia Y. Suto & 

Suyama 2005 (H) 

Myrioconium Syd. & P. Syd. 1912, 

anamorphic Ciboria Fuckel 1870, 

Ciborinia Whetzel 1945, (Fr.) P. Karst. 

1871, Martininia Dumont & Korf 1970, 

Myriosclerotinia N.F. Buchw. 1947, 

Sclerotinia Fuckel 1870 Stromatinia 

(Boud.) Boud. 1907 and Monilinia 

Honey 1928 (H) 

Ovulitis N.F. Buchw. 1970, anamorphic 

Ovulinia F.A. Weiss 1940 (H) 

Sclerotium Tode 1790, anamorphic Myriosclerotinia 

N.F. Buchw. 1947, Sclerotinia 

Fuckel 1870 and Stromatinia 

(Boud.) Boud. 1907 (H) 

Septotis N.F. Buchw. ex Arx 1970, 

anamorphic Septotinia Whetzel ex J.W. 

Groves & M.E. Elliott 1961 (H) 

Streptobotrys Hennebert 1973, anamorphic 

Streptotinia Whetzel 1945 (H) 

Valdensia Peyronel 1923, anamorphic 

Valdensinia Peyronel 1953 (H) 


Verrucobotrys Hennebert 1973, anamorphic 

Seaverinia Whetzel 1945 (H) 

Vibrisseaceae Korf 

Acephala Grünig & T.N. Sieber 2005 (H) 

Anavirga B. Sutton 1975, anamorphic 

Vibrissea Fr. 1822 (H) 

Phialocephala W.B. Kendr. 1961, some 

possibly anamorphic Vibrissea Fr. 1822 

(H) 

Phialophora-like, anamorphic Chlorovibrissea 

L.M. Kohn 1989 (H) 

Helotiales, genera incertae sedis 

Catenulifera Hosoya 2002, anamorphic 

Hyphodiscus Kirschst. 1906 (H) 

Chalara (Corda) Rabenh. 1844 (H)* 

Clathrosporium Nawawi & Kuthub. 1987 

(H) 

Crucellisporium M.L. Farr 1968 (C) 

Cystotricha Berk. & Broome 1850 (C) 

Dactylaria Sacc. 1880 (H) 

Diplococcium Grove 1885 (H) 

Endoconidium Prill. & Delacr. 1891, anamorphic 

Gloeotinia M. Wilson, Noble 

& E.G. Gray 1954 (H) 

Fontanospora Dyko 1978 (H) 

Fulvoflamma Crous 2006 (H) 

Glarea Bills & Paláez 1999 (H) 

Helgardia Crous & W. Gams 2003, anamorphic 

Oculimacula Crous & W. 

Gams 2003 (H) 

Humicolopsis Cabral & S. Marchand 1976 

(H) 

Hyalodendriella Crous 2007 (H) 

Hysteropezizella Höhn. 1917 (C) 

Lemonniera De Wild. 1894 (H) 

Leptodontidium de Hoog 1979 (H) 

Libartania Nag Raj 1979 (C) 

Margaritispora Ingold 1942 (H) 

Meliniomyces Hambl. & Sigler 2005 (H) 

Mirandina G. Arnaud ex Matsush. 1975 

(H) 

Monochaetiellopsis B. Sutton & DiCosmo 

1977, anamorphic Hypnotheca Tommerup 

1970 (C) 

Mycochaetophora Hara & Ogawa 1931 

(H) 

Neozythia Petr. 1958 (C) 

Rhexocercosporidium U. Braun 1994 (H) 

Rhizocladosporium Crous & U. Braun

2007 (H) 

Rhodesia Grove 1937 (C) 

Scytalidium Pesante 1957 (H) 

Sphaerographium Sacc. 1884 (C) 

Spirosphaera Beverw. 1953 (H) 

Tetracladium De Wild. 1893 (H) 

Thedgonia B. Sutton 1973 (H) 

Trimmatostroma Corda 1837 (H) 

Variocladium Descals & Marvanová 1999 

(H) 

Xenopolyscytalum Crous 2011(‘2010’) (H) 

Rhytismatales M.E. Barr ex Minter 

Ascodichaenaceae D. Hawksw. & Sherwood 

Dichaenopsella Petr. 1952, anamorphic 

Ascodichaena Butin 1977 (C) 

Macroallantina Speer 1987, anamorphic 

Delpinoina Kuntze, 1891 (C) 

Polymorphum Chevall. 1822, anamorphic 

Ascodichaena Butin 1977 (C) 

Cryptomycetaceae Höhn. 

?Phacidiopycnis Potebnia 1912, anamorphic 

Potebniamyces Smerlis 1962 (C) 

Rhytismataceae Chevall. 

Conostroma Moesz 1921, anamorphic 

Colpoma Wallr. 1833 (C) 

Crandallia Ellis & Sacc. 1897, anamorphic 

Bifusella Höhn. 1917 and 

Duplicaria Fuckel 1870 (C) 

Cryocaligula Minter 1986, anamorphic 

Ploioderma Darker. 1967 (C) 

Hysterodiscula Petr. 1942, anamorphic 

Duplicaria Fuckel 1870 (C) 

Leptostroma Fr. 1815, anamorphic Lophodermium 

Chevall. 1826 (C) 

Melasmia Lév. 1846, anamorphic Rhytisma 

Fr. 1818 (C) 

Tryblidiopycnis Höhn. 1918, anamorphic 

Tryblidiopsis P. Karst. 1871 (C) 

Rhytismatales, genera incertae sedis 

Fuligomyces Morgan-Jones & Kamal 1984 

(H) 

Uyucamyces H.C. Evans & Minter 1985, 

anamorphic Ocotomyces H.C. Evans & 

Minter 1985 (C) 

Leotiomycetes, families incertae sedis 

Myxotrichaceae Currah 


Geomyces Traaen 1914, anamorphic 

Pseudogymnoascus Raillo 1929 (H) 

?Gymnostellatospora Udagawa, Uchiy. & 

Kamiya 1993, anamorphic Pseudogymnoascus 

Raillo 1929 (C) 

?Malbranchea Sacc. 1882, anamorphic 

Myxotrichum Kunze 1823 (H) 

Oidiodendron Robak 1932, anamorphic 

Byssoascus Arx 1971 and Myxotrichum 


Leotiomycetes, genera incertae sedis 

Brachyalara Réblová & W. Gams 2011 

(H) 

Chaetomella Fuckel 1870 (C) 

Collophora Damm & Crous 2010 (C) 

Eleutheromyces Fuckel 1870 (C) 

Geniculospora Sv. Nilsson ex Marvanová 

& Sv. Nilsson 1971 (H) 

Hyphozyma de Hoog & M.T. Sm. 1981 

(H) 

Infundichalara Réblová & W. Gams 2011 

(H) 

Leohumicola N.L. Nick. 2005 (H) 

Meliniomyces Hambl. & Sigler 2005 (H) 

Scytalidium Pesante 1957, anamorphic 

Xylogone Arx & T. Nilsson 1969 (H) 

Tetrachaetum Ingold 1942 (H) 

Tiarosporella Höhn. 1924, anamorphic 

Darkera H.S. Whitney, J. Reid & Piroz. 

1975 (C) 

Class Orbiliomycetes O.E. Erikss. & Baral 

Orbiliales Baral, O.E. Erikss., G. Marson & E. 

Weber 

Orbiliaceae Nannf. 


anamorphic Orbilia Fr. 1836 (H) 

Arthrobotrys Corda 1939, anamorphic 

Orbilia Fr. 1836 (H) 

Brachyphoris Juan Chen, L.L. Xu, B. Liu 

& Xing Z. Liu 2007, anamorphic 

Hyalorbilia Baral & G. Marson 2000 

(H) 

Dactylaria Sacc. 1880, anamorphic Orbilia 

Fr. 1836 (H) 

Dactylella Grove 1884, anamorphic Orbilia 

Fr. 1836 (H) 

Dactylellina M. Morelet 1968 (H) 

Dicranidion Harkn. 1885, anamorphic Orbilia 

Fr. 1836 (H) 

179

180 

Drechslerella Subram. 1964, anamorphic 


Dwayaangam Subram. 1978, anamorphic 


Gamsylella M. Scholler, Hagedorn & A. 

Rubner 1999, anamorphic Orbilia Fr. 

1836 (H) 

Helicoön Morgan 1892, anamorphic 



anamorphic Orbilia Fr. 1836 (H) 

Monacrosporium Oudem. 1885, anamorphic 


Pseudotripoconidium Z.F. Yu & K.Q. 

Zhang 2011, anamorphic Orbilia Fr. 

1836 (H)* 

Tricellula Beverw. 1954, anamorphic 


Tridentaria Preuss 1852 (H) 

Trinacrium Riess 1852, anamorphic Orbilia 

Fr. 1836 (H) 

Class Pezizomycetes sensu O.E. Erikss. & 

Winka 

Pezizales J. Schröt. 

Ascobolaceae Boud. ex Sacc. 


Ascobolus Pers. 1796 (H) 

Papulospora-like, anamorphic Cubonia 

Sacc. 1889 (H) 

Rhizostilbella Wolk 1914, anamorphic 

Ascobolus Pers. 1792 (H) 

Caloscyphaceae Harmaja 

Geniculodendron G.A. Salt 1974, anamorphic 

Caloscypha Boud. 1885 (H) 

Chorioactidaceae Pfister 

Kumanasamuha P.Rag. Rao & D. Rao 

1964, anamorphic Chorioactis Kupfer 

ex Eckblad 1968 (H) 

Verticicladium Preuss 1851, anamorphic 

Desmazierella Lib. 1829 (H) 

Morchellaceae Reichenb. 

Costantinella Matr. 1892, anamorphic 

Morchella Dill. ex Pers. 1794 (H) 

Pezizaceae Dumort. 

Chromelosporium Corda 1833, anamor- 


phic Muciturbo P.H.B. Talbot 1989, 

Peziza Dill. ex Fr. 1822 and Plicaria 

Fuckel 1870 (H) 

Glischroderma Fuckel 1870 (C) 

Oedocephalum Preuss 1851, anamorphic 

Iodophanus Korf 1967 and Peziza Dill. 

ex Fr. 1822 (H) 

Ostracoderma Fr. 1825, anamorphic Peziza 

Dill. ex Fr. 1822 (C) 

Pyronemataceae Corda 

Actinosporella Descals, Marvanová & J. 

Webster 1999, anamorphic Miladina 

Svrček 1972 (H) 

Ascorhizoctonia Chin S. Yang & Korf 

1985, anamorphic Tricharina Eckblad 

1968 (H) 

Basifimbria Subram. & Lodha 1968, 

anamorphic Geopyxis (Pers.) Sacc. (H) 

Complexipes C. Walker 1979, anamorphic 

Tricharina Eckblad 1968 and Wilcoxina 

Chin S. Yang & Korf 1985 (H) 

?Dichobotrys Hennebert 1973, anamorphic 

Pyropyxis Egger 1984 and Trichophaea 

Boud. 1885 (H) 

Micronematobotrys Xiang Sun & L.D. 

Guo 2010 (H) 

?Molliardiomyces Paden 1984, anamorphic 

Stephensia Tul. & C. Tul. 1851 

(H) 

?Scytalidium Pesante 1957, anamorph of 

Anthracobia Boud. 1885 (H) 

Rhizinaceae Bonord. 

Phymatotrichopsis Hennebert 1973 (H) 

Sarcoscyphaceae LeGal ex Eckblad 

Molliardiomyces Paden 1984, anamorphic 

Nanoscypha Denison 1972, Phillipsia 

Berk. 1881, Pithya Fuckel 1870 and 

Sarcoscypha (Fr.) Boud. 1885 (H) 

Verticicladium Preuss 1851, anamorphic 

Desmazierella Lib. 1829 (H) 

Sarcosomataceae Kobayasi 

Conoplea Pers. 1797, anamorphic Urnula 

Fr. 1849 and Plectania Fuckel 1870 (H) 

Strumella Fr. 1849, anamorphic Urnula Fr. 

1849 (H) 

Verticicladium Preuss 1851, anamorphic

Sarcosoma Casp. 1891 (H) 

Pezizales, genera incertae sedis 

Cephaliophora Thaxt. 1903 (H) 

Class Sordariomycetes sensu O.E. Erikss. & 

Winka 

Subclass Hypocreomycetidae O.E. Erikss. & 

Winka 

Coronophorales Nannf. 

Chaetosphaerellaceae Huhndorf, A.N. Mill. & 

F.A. Fernández 

Oedemium Link 1824, anamorphic Chaetosphaerella 

E. Müll. & C. Booth 1972 

(H) 

Veramycina Subram. 1995, anamorphic 

Chaetosphaerella E. Müll. & C. Booth 

1972 (H) 

Nitschkiaceae (Fitzp.) Nannf. 

Acremonium-like, anamorphic Acanthonitschkea 

Speg. (H) 

Glomerellales Chadef. ex Réblová, W. Gams 

& Seifert 

Australiascaceae Réblová & W. Gams 

Monilochaetes Halst. ex Harter 1916, 

anamorphic Australiasca Sivan. & 

Alcorn 2002 (H)* 

Glomerellaceae Locq. ex Seifert & W. Gams 

Colletotrichum Corda 1831, anamorphic 

Glomerella Spauld. & Schrenk 1903 

(C) 

Reticulascaceae Réblová & W. Gams 

Cylindrotrichum Bonord.1851, anamorphic 

Reticulascus Réblová & W. Gams 

2011 (H)* 

Hypocreales Lindau 

Bionectriaceae Samuels & Rossman 

Acremonium-like, anamorphic Dimerosporiella 

Speg. 1908, Ijuhya Starbäck 

1899, Hydropisphaera Dumont 1822, 

Lasionectria (Sacc.) Cooke 1884, Mycoarachis 

Malloch & Cain 1970, 

Mycocitrus Moller 1901, Nectriella 

Nitschke ex Fuckel 1870, Nectriopsis 

Maire 1911, Nigrosabulum Malloch & 

Cain 1970, Protocreopsis Yoshim. Doi 

1977, Selinia P. Karst. 1876, and 

Stilbocrea Pat. 1900 and Verrucostoma 


Hirooka, Tak. Kobay. & Chaverri 2010 

(H) 

Albosynnema E.F. Morris 1967 (H) 

Clonostachys Corda 1839, anamorphic 

Bionectria Speg. 1919 (H) 

Dendrodochium-like, anamorphic Nectriella 


Didymostilbe Henn. 1902, anamorphic 

Peethambara Subram. & Bhat 1978 (H) 

Geosmithia J.Pitt 1979 (H)* 

Gliocladium-like, anamorphic Roumegueriella 

Speg. 1880 (H) 

Gliomastix Guég. 1905 (H)* 

Gracilistilbella Seifert 2000, anamorphic 

Stilbocrea Pat. 1900 (H) 

Kutilakesa Subram. 1956, anamorphic 

Nectriella Nitschke ex Fuckel 1870 (H) 

Rhopalocladium Schroers, Samuels & W. 

Gams 1999, anamorphic Nectriopsis 

Maire 1911 (H) 

Sarocladium W. Gams & D. Hawksw. 

1976 (H) 

Spicellum Nicot & Roquebert 1976 (H) 

Stanjemonium W. Gams, O'Donnell, 

Schroers & M. Chr. 1999 (H) 

Stilbella Lindau 1900 anamorphic 

Stilbocrea Pat. 1900 (H) 

Trichothecium-like, anamorphic Heleococcum 

C.A. Jørg. 1922 (H) 

Tubercularia-like, anamorphic Selinia P. 

Karst. 1876 (H) 

Verticillium-like, anamorphic Nectriopsis 

Maire 1911 (H) 

Vesicladiella Crous & M.J. Wingf. 1994 

(H) 

Virgatospora Finley 1967, anamorphic 


Clavicipitaceae (Lindau) Earle ex Rogerson 

Acremonium-like, anamorphic Neobarya 

Lowen 1986 (H) 

Albomyces I. Miyake 1908, anamorphic 

Aciculosporium I. Miyake (C) 

Amphichordia Fr., 1825 (H) 

Aschersonia Mont. 1848, anamorphic 

Hypocrella Sacc. and Moelleriella Bres. 

1897 (C) 

Blistum B. Sutton 1973, anamorphic 

Berkelella (Sacc.) Sacc. 1891 (H) 

Calcarisporium Preuss 1851, anamorphic 

Neobarya Lowen (H) 

Corallocytostroma Y.N. Yu & Z.Y. Zhang 

181

182 

1980 (C) 

Diploöspora Grove 1916, anamorphic 

Neobarya Lowen 1986 (H) 

Drechmeria W. Gams & H.-B. Jansson 

1985 (H) 

Ephelis Fr. 1849, anamorphic Atkinsonella 

Diehl, Balansia Speg., 1885, Epichloë 

(Fr.) Tul. & C. Tul., 1865, Dothichloë 

G.F. Atk. 1894, Heteroepichloë E. 

Tanaka, C. Tanaka, Gafur & Tsuda 

2002, Nigrocornus Ryley & Langdon 

2003 and Myriogenospora G.F. Atk. 

1894 (H) 

Harposporium Lohde 1874, anamorphic 

Podocrella Seaver 1928 (H) 

Hirsutella-like, anamorphic Orbiocrella D. 

Johnson, G.H. Sung, Hywel-Jones & 

Spatafora 2009 and Moelleriella Bres. 

1897 (H) 

Lecanicillium W. Gams & Zare 2001, 

anamorphic Hypocrella Sacc. 1878and 

Neobarya Lowen 1986 (H) 

Metarhizium Sorokīn 1879, anamorphic 

Metacordyceps G.H. Sung, J.M. Sung, 

Hywel-Jones & Spatafora 2007 (H) 

?Munkia Speg. 1886, anamorphic Mycomalus 

A. Møller 1901 (H) 

Neomunkia Petr. 1947 (H) 

Neotyphodium Glenn, C.W. Bacon & 

Hanlin 1996, anamorphic Epichloë (Fr.) 

Tul. & C. Tul. 1865 (H) 

Nomuraea Maubl. 1903 (H) 

Pleurodesmospora Samson, W. Gams & 

H.C. Evans 1979 (H) 

Pochonia Bat. & O.M. Fonseca 1965 (H) 

Polycephalomyces Kobayasi 1941, anamorphic 

Berkelella (Sacc.) Sacc. 1891 

(H) 

Pseudogibellula Samson & H.C. Evans 

1973, anamorphic Torrubiella Boud. 

1885 (H) 

Pseudomeria G.L. Barron 1980 (H) 

Rotiferophthora G.L. Barron 1991 (H) 

Sphacelia Lév. 1827, anamorphic Atkinsonella 

Diehl 1950 and Claviceps Tul. 

1853 (H) 

Ustilaginoidea Bref. 1895, anamorphic 

Villosiclava E. Tanaka & C. Tanaka 

2009 (H) 

Cordycipitaceae Kreisel ex G.M. Sung, J.M. 

Sung, Hywel-Jones & Spatafora 


Akanthomyces Lebert 1858, anamorphic 

Cordyceps Fr. 1824 and Torrubiella 

Boud. 1885 (H) 

Beauveria Vuill. 1912, anamorphic Cordyceps 

Fr. 1824 (H)* 

Engyodontium de Hoog 1978 (H) 

Gibellula Cavara 1894, anamorphic 

Cordyceps Fr. 1824 and Torrubiella 

Boud. 1885 (H) 

Granulomanus de Hoog & Samson 1978, 

anamorphic Torrubiella Boud. 1885 

(H) 

Hirsutella Pat. 1892, anamorphic 

Cordyceps Fr. 1824 and Cordycepioideus 

Stifler 1941 (H) 

Isaria Pers. 1794, anamorphic Cordyceps 

Fr. 1824 and Torrubiella Boud. 1885 

(H) 

Lecanicillium W. Gams & Zare 2001, 

anamorphic Cordyceps Fr. 1824 and 

Torrubiella Boud. 1885 (H) 

Microhilum H.Y. Yip & A.C. Rath 1989 

(H) 

Pseudogibellula Samson & H.C. Evans 

1973 (H) 

Rotiferophthora G.L. Barron 1991 (H) 

Septofusidium W. Gams 1971, anamorphic 

Cordyceps Fr. 1824 (H) 

Simplicillium W. Gams & Zare 2001, 

possibly anamorphic Torrubiella Boud. 

1885 (H) 

Tolypocladium W. Gams 1971, anamorphic 

Cordyceps Fr. 1824 (H) 

Hypocreaceae De Not. 

Acremonium-like, anamorphic Protocrea 

Petch 1937 and Pseudohypocrea 

Yoshim. Doi 1972 (H) 

Arachnocrea Z. Moravec 1956, anamorphic 

Protocrea Petch 1937 and 

Sarawakus Lloyd 1924 (C) 

Cladobotryum Nees 1816, anamorphic 

Hypomyces (Fr.) Tul. & C. Tul. 1860 

(H)* 

Gliocladium Corda 1840, anamorphic 

Hypocrea Fr. 1825 and Sphaerostilbella 

(Henn.) Sacc. & D. Sacc. 1905 

(H) 

Gliocladium-like, anamorphic Sarawakus 


Mycogone Link 1809, anamorphic Hypomyces 

(Fr.) Tul. & C. Tul. 1860 (H)

Papulaspora-like, anamorphic Hypomyces 

(Fr.) Tul. & C. Tul. 1860 (H) 

Sepedonium Link 1809, anamorphic Hypomyces 

(Fr.) Tul. & C. Tul. 1860 (H) 

Sibirina G.R.W. Arnold 1970, anamorphic 


(H) 

Stephanoma Wallr. 1833, anamorphic 


(H) 

Stilbella Lindau 1900, anamorphic 

Hypocrea Fr. 1825 (H) 

Stromatocrea W.B. Cooke 1952, anamorphic 

Hypocreopsis P. Karst. 1873 (H) 

Trichoderma Pers. 1794, anamorphic 

Hypocrea Fr. 1825 and Podostroma P. 

Karst. 1892 (H)* 

Trichoderma-like, anamorphic Sarawakus 


Trichothecium-like, anamorphic Hypomyces 

(Fr.) Tul. & C. Tul. 1860 (H) 

Verticillium-like, anamorphic Aphysiostroma 

Barrasa, A.T. Martínez & G. 

Moreno 1986, Sphaerostilbella Henn. 

1901 and Coniochaeta (Sacc.) Cooke 

1887 (H) 

Verticimonosporium Matsush. 1971 (H) 

Nectriaceae Tul. & C. Tul. 

Acremonium-like, anamorphic Cosmospora 

Rabenh. 1862, Nectria (Fr.) Fr. 1849 

and Neocosmospora E.F. Sm. 1899 (H) 

Actinostilbe Petch 1925, anamorphic Lanatonectria 

Samuels & Rossman 1999 

(H) 

Antipodium Piroz. 1974, anamorphic 

Ophionectria Sacc. 1878 (H) 

Aphanocladium W. Gams 1971 (H) 

Atractium Link 1809 (H) 

Calostilbella Höhn. 1919, anamorphic 

Calostilbe Sacc. & Syd. 1902 (H) 

Campylocarpon Halleen, Schroers & 


Chaetodochium Höhn. 1932, anamorphic 

Pseudonectria Seaver 1909 (H) 

Chaetopsina Rambelli 1956, anamorphic 

Chaetopsinectria J. Luo & W.Y. 

Zhuang and Cosmospora Rabenh. 1862 

(H) 

Curvicladiella Decock & Crous 2006 (H) 

Curvicladium Decock & Crous 1998 (H) 


Cyanochyta Höhn. 1915, anamorphic 

Gibberella Sacc. 1877 (C) 

Cyanophomella Höhn. 1918, anamorphic 


Cylindrocarpon Wollenw. 1913, anamorphic 

Neonectria Wollenw. 1917, 

Rugonectria P.Chaverri & Samuels 

2011, Thelonectria P. Chaverri & C.G. 

Salgado 2011, Ilyonectria P.Chaverri & 

Samuels 2011 (H)* 

Cylindrocladiella Boesew. 1982, anamorphic 

Cosmospora Rabenh. 1862 and 

Nectricladiella Crous & C.L. Schoch 

2000 (H) 

Cylindrocladium Morgan 1892, anamorphic 

Calonectria De Not. 1867 (H) 

Dacryoma Samuels 1988, anamorphic 

Nectria (Fr.) Fr. 1849 (H) 

Dematiocladium Allegr., Aramb., Cazau & 


Dichomera Cooke 1878, anamorphic 


Flagellospora Ingold 1942, anamorphic 

Nectria (Fr.) Fr. 1849 (H) 

Fusarium Link 1809, anamorphic Albonectria 

Rossman & Samuels 1999, Cyanonectria 

Samuels & Chaverri 2009, 

Gibberella Sacc. 1877, Corallomycetella 

Henn. 1904 and Haematonectria 

Samuels & Nirenberg 1999 and 

unknown teleomorphs (H)* 

Fusarium-like, anamorphic, Geejayessia 

Schroers, Gräfenhan & Seifert 2011 and 

Cyanonectria Samuels & Chaverri 2009 

(H)* 

Fusicolla Bonord. 1851 (H) 

Gliocephalotrichum J.J. Ellis & Hesselt. 

1962, anamorphic Leuconectria Rossman, 

Samuels & Lowen 1993 (H) 

Gliocladiopsis S.B. Saksena 1954, anamorphic 

Glionectria Crous & S.L. 

Schoch 2000 (H) 

Gyrostroma Naumov 1914, anamorphic 

Nectria (Fr.) Fr. 1849 (C) 

Heliscus Sacc. 1880, anamorphic Neonectria 

Wollenw. 1917 . 1849 (H) 

Macroconia (Wollenw.) Gräfenhan, 

Seifert & Schroers 2011 (H) 

Mariannaea G. Arnaud ex Samson 1974 

anamorphic ‘Nectria’ (Fr.) Fr. 1849 and 

Cordyceps Fr. 1833 (H)* 

183

184 

Microcera Desm. 1848, anamorphic 

Gibberella Sacc. 1877 (H)* 


anamorphic Chaetonectrioides Matsush. 

(H) 

Nalanthamala Subram. 1956, anamorphic 

Rubrinectria Rossman & Samuels 1999 

(H) 

Penicillifer Emden 1968, anamorphic Viridispora 

Samuels & Rossman 1999 (H) 

Pleurocolla Petr. 1924 (H) 

Rhizostilbela Wolk 1914, anamorphic 

Corallomycetella Henn. 1904 (H) 

Septofusidium W. Gams 1971 (H) 

Stachybotryna Tubaki & T. Yokoy. 1971, 

anamorphic ‘Nectria’ (Fr.) Fr. 1849 (H) 

Stilbella Lindau 1900, anamorphic Cosmospora 

Rabenh. 1862 and Nectria (Fr.) 

Fr. 1849 (H)* 

Tilachlidium Preuss 1851, anamorphic 

‘Pseudonectria’ Seaver 1909 (H) 

Tubercularia Tode 1790, anamorphic 

Nectria (Fr.) Fr. 1849 (H)* 

Verticillium-like, anamorphic Cosmospora 

Rabenh. 1862 (H) 

Volutella Fr. 1832, anamorphic Cosmospora 

Rabenh. 1862 and Pseudonectria 

Seaver 1909 (H) 

Xenocylindrocladium Decock, Hennebert 

& Crous 1997, anamorphic Xenocalonectria 

Crous & C.L. Schoch 2000 (H) 

Zythiostroma Höhn. ex Falck 1923, 

anamorphic Nectria (Fr.) Fr. 1849 (C) 

Niessliaceae Kirschst. 

Acremonium-like, anamorphic Trichosphaerella 

E. Bommer, M. Rousseau & 

Sacc. 1891 and Valetoniellopsis Samuels 

& M.E. Barr 1998 (H) 

Custingophora-like, anamorphic Melanopsamma 

Niessl 1876 (H) 

Monocillium S.B. Saksena 1955, anamorphic 

Hyaloseta A.W. Ramaley 2001 

and Niesslia Auersw. 1869 (H) 

Ophiocordycipitaceae G.H. Sung, J.M. Sung, 

Hywel-Jones & Spatafora 

Chaunopycnis W. Gams 1979 (H) 

Haptocillium W. Gams & Zare 2001 

anamorphic Ophiocordyceps Petch 

1931 (H) 


Hirsutella Pat. 1892, anamorphic Ophiocordyceps 

Petch 1931 (H) 

Hymenostilbe Petch 1931, anamorphic 

Ophiocordyceps Petch 1931 (H) 

Paraisaria Samson & B.L. Brady 1983, 

anamorphic Ophiocordyceps Petch 

1931 (H) 

Purpureocillium Luangsa-ard, Hywel- 

Jones, Houbraken & Samson 2011 (H) 

Syngliocladium Petch 1932, anamorphic 

Ophiocordyceps Petch 1931(H) 

Tolypocladium W. Gams 1971, anamorphic 

Elaphocordyceps G.H. Sung & 

Spatafora 2007 (H) 

Verticillium-like, anamorphic Elaphocordyceps 

G.H. Sung & Spatafora 2007 

(H) 

Hypocreales, genera incertae sedis 

Acremonium Link 1809 (H)* 

Acremonium-like, anamorphic Emericellopsis 

J.F.H. Beyma 1940 and Leucosphaerina 

Arx 1987 (H) 

Ascochytopsis-like, anamorphic Eucasphaeria 


Cancellidium Tubaki 1975 (H) 

Didymostilbe Henn. 1902, anamorphic 


Escovopsis J.J. Muchovej & Della Lucia 

1990 (H) 

Hapsidospora Malloch & Cain 1970 (C) 

Haptospora G.L. Barron 1991 (H) 

Illosporiopsis D. Hawksw. 2001 (H) 

Illosporium Mart. 1817 (H) 

Imicles Shoemaker & Hambl. 2001 (H) 

Memnoniella Höhn. 1923 (H) 

Myrothecium Tode 1790 (H) 

Parasarcopodium Melnik, S.J. Lee & 


Pseudomicrodochium B. Sutton 1975 (H) 

Sarocladium W. Gams & D. Hawksw. 

1976 (H) 

Septomyrothecium Matsush. 1971, anamorphic 

Peethambara Subram. & Bhat 

1978 (H) 

Sorosporella Sorokīn 1888 (H) 

Spicellum Nicot & Roquebert 1976 (H) 

Sporothrix Hektoen & C.F. Perkins 1901, 

anamorphic Leucosphaerina Arx 1987 

(H) 

Stachybotrys Corda 1837 (H)

Stilbella Lindau 1900, anamorphic Emericellopsis 

J.F.H. Beyma 1940 and 

Valsonectria Speg. 1881 (H) 

Tilachlidium Preuss 1851 (H) 

Trichothecium Link 1809 (H)* 

Lulworthiales Kohlm., Spatafora & Volkm.- 

Kohlm. 2000 

Lulworthiaceae Kohlm., Spatafora & Volkm.- 

Kohlm. 2000 


Lindra I.M. Wilson 1956 (H) 

Melanosporales N. Zhang & M. Blackw. 

Ceratostomataceae G. Winter 

Acrospeira Berk. & Broome 1857 (H) 

Gonatobotrys Corda 1839, anamorphic 

Melanospora Corda 1837 (H) 

Harzia Costantin 1888, anamorphic Melanospora 

Corda 1837 (H) 

Papulaspora Preuss 1851, anamorphic 

Melanospora Corda 1837 and Microthecium 

Corda 1842 (H) 

Proteophiala Cif. 1958, anamorphic Melanospora 

Corda 1837, Persiciospora 

P.F. Cannon & D. Hawksw. 1982, 

Sphaerodes Clem. 1909, Pteridiosperma 

J.C. Krug & Jeng 1979 (H) 

[Sphaerodes Clem. 1909 is a teleomorph 

which was reported to produce ampulliform 

phialides on irregularly branched 

conidiophores diectrly on ascomata or 

on surrounding hyphae (Vujanovic & 

Goh, 2009)] 

Microascales Luttr. ex Benny & Kimbr. 

Chadefaudiellaceae Faurel & Schotter ex 

Benny & Kimbr. 

Arthrographis-like, anamorphic Faurelina 

Locq.-Lin. 1975 (H) 

Gondwanamycetaceae Réblová, W. Gams & 

Seifert 

Custingophora Stolk, Hennebert & 

Klopotek 1968, anamorphic Gondwanamyces 

G.J. Marais & M.J. Wingf. 1998 

(H)* 

Halosphaeriaceae E. Müll. & Arx ex Kohlm. 

Cirrenalia Meyers & R.T. Moore 1960 

(H) 


Clavariopsis-like, anamorphic Corollospora 

Werderm. 1922 (H) 

Clavatospora Sv. Nilsson ex Marvanová 

& Sv. Nilsson 1971, anamorphic Corollospora 


Culcitalna Meyers & R.T. Moore 1960, 

anamorphic Halosarpheia Kohlm. & E. 

Kohlm. 1977 (H) 

Halosigmoidea Nakagiri, K.L. Pang & 

E.B.G. Jones 2009, anamorphic Corollospora 


Humicola Traaen 1914, anamorphic Pseudolignincola 

Chatmala & E.B.G. Jones 

2006 and Lignincola Höhnk 1955 (H) 


Nereiospora E.B.G. Jones, R.G. Johnson 

& S.T. Moss 1983 (H) 

Periconia Tode 1791, anamorphic Halosphaeria 

Linder 1944 (H) 

Sigmoidea J.L. Crane 1968, anamorphic 

Corollospora Werderm. 1922 (H) 

Trichocladium Harz 1871, anamorphic 

Halosphaeria Linder 1944 (H) 

Varicosporina Meyers & Kohlm. 1965, 

anamorphic Corollospora Werderm. 

1922 (H) 

Microascaceae Luttr. ex Malloch 

Atrhrographis G. Cochet ex Sigler & J.W. 

Carmich. 1976, anamorphic Canariomyces 

Arx (H) 

Brachyconidiellopsis Decock, R.F. Castañeda 

& Adhikari 2004 (H) 

Cephalotrichum Link 1809, anamorphic 

Microascus Zukal 1885 (H) 

Echinobotryum Corda 1831 (H) 

Graphium Corda 1837, anamorphic Microascus 

Zukal 1885, Petriella Curzi 1930 

and Pseudallescheria Negr. & I. Fisch. 

1944 (H) 

Humicola Traaen 1914, anamorphic Lophotrichus 

R.K. Benj. 1949 (H) 

Parascedosporium Gilgado, Gené, Cano & 

Guarro 2007 (H) 

Scedosporium Sacc. ex Castell. & Chalm. 

1919, anamorphic Pseudallescheria 

Negr. & I. Fisch. 1944 and possibly 

Petriella Curzi 1930 (H) 

Scopulariopsis Bainier 1907, anamorphic 

Microascus Zukal 1885 and ?Petriella 

Curzi 1930 and Pithoascus Arx 1973 

(H) 

185

186 

Trichurus Clem. 1896 (H) 

Wardomyces F.T. Brooks & Hansf. 1923, 

anamorphic Microascus Zukal 188 (H) 

Wardomycopsis Udagawa & Furuya 1978, 

anamorphic Microascus Zukal 1885 

(H) 

Microascales, genera incertae sedis 

Bisporostilbella Brandsb. & E.F. Morris 

1971 (H) 

Chalara-like, anamorphic Ceratocystis 

Ellis & Halst. 1890 and Cornuvesica 

C.D. Viljoen, M.J. Wingf. & K. Jacobs 

2000 (H) 

Gabarnaudia Samson & W. Gams 1974, 

anamorphic Sphaeronaemella P. Karst. 

1884 (H) 

Sporendocladia G. Arnaud ex Nag Raj & 

W.B. Kendr. 1975 (H) 

Thielaviopsis Went 1893, anamorphic 

Ceratocystis Ellis & Halst. 1890 (H) 

Hypocreomycetidae, families incertae sedis 

Plectosphaerellaceae W. Gams, Summerbell 

& Zare 

Acremonium Link 1809 (H)* 

Acrostalagmus Corda 1838 (H)* 

Gibellulopsis Bat. & H. Maia 1959 (H) 

Musicillium Zare & W. Gams 2007 (H) 

Plectosporium M.E. Palm, W. Gams & 

Nirenberg 1995, anamorphic Plectosphaerella 

Kleb. 1929 (H) 

Stachylidium Link 1809 (H)* 

Verticillium Nees 1816 (H) 

Hypocreomycetidae, genera incertae sedis 

Canalisporium Nawawi & Kuthub. 1989, 

anamorphic Ascothailandia Sriindrasutdhi, 

Boonyuen, Sivichai & E.B.G. 

Jones 2010 (H) 

Dictyochaeta Speg. 1923, anamorphic 

Ascocodinaea Samuels, Candoussau & 

Magni 1997 (H) 

?Glomerulispora Abdel-Wahab, Abdel- 

Aziz & Nagahama 2010, anamorphic 

Torpedospora Meyers 1957 (H) 

Moheitospora Abdel-Wahab, Abdel-Aziz 

& Nagahama 2010, anamorphic Juncigena 

Kohlm., Volkm.-Kohlm & O.E. 

Erikss. 1997 (H) 


Sporoschismopsis Hol.-Jech. & Hennebert 

1972, anamorphic Porosphaerellopsis 

Samuels & E. Müll. 1982 (H) 

Subclass Sordariomycetidae O.E. Erikss. & 

Winka 

Boliniales P.F. Cannon 2001 

Boliniaceae Rick 1931 

Phaeoisaria Höhn. 1909 anamorphic 

Lentomitella Höhn. 1906 (H) 

Calosphaeriales M.E. Barr 

Calosphaeriaceae Munk 

Calosphaeriophora Réblová, L. Mostert, 

W. Gams & Crous 2004, anamorphic 

Calosphaeria Tul. & C. Tul. (H) 

Lecythophora Nannf. 1934, anamorphic 

Sulcatistroma A.W. Ramaley 2005 (H) 

Phaeocrella Réblová, L. Mostert, W. 

Gams & Crous 2004, anamorphic Togniniella 

Réblová, L. Mostert, W. Gams 


Phialophora-like, anamorphic Jattaea 

Berl. 1900 (H) 

Stachybotrys-like, anamorphic Jattaea 

Berl. 1900 (H) 

Pleurostomataceae Réblová, L. Mostert, W. 

Gams & Crous 

Pleurostomophora Vijaykr., L. Mostert, 

Jeewon, W. Gams, K.D. Hyde & Crous 

2004, anamorphic Pleurostoma Tul. & 

C. Tul. 1863 (H) 

Chaetosphaeriales Huhndorf, A.N. Mill. & 

F.A. Fernández 

Chaetosphaeriaceae Réblová, M.E. Barr & 

Samuels 

Cacumisporium Preuss 1851, anamorphic 

Chaetosphaeria Tul. & C. Tul. 1863 

(H) 

Catenularia Grove 1886, anamorphic 


(H) 


Ascochalara Réblová 1999 and 


(H)* 

Chloridium Link 1809, anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 and

Melanopsammella Höhn. 1920 (H) 

Codinaea Maire 1937, anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

?Cordana Preuss 1851, anamorphic Porosphaerella 

E. Müll. & Samuels 1982 

(H) 

Craspedodidymum Hol.-Jech. 1972, anamorphic 

Chaetosphaeria Tul. & C. Tul. 

1863 (H) 

Cryptophiale Piroz. 1968, anamorphic 


(H) 

Custingophora Stolk, Hennebert & Klopotek 

1968 anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

Dictyochaeta Speg. 1923, anamorphic 


and Striatosphaeria Samuels & E. Müll. 

1979 (H) 

Exserticlava S. Hughes 1978, anamorphic 


(H) 

Fusichalara S. Hughes & Nag Raj 1973, 

anamorphic Chaetosphaeria Tul. & C. 

Tul. 1863 (H) 

Gonytrichum Nees & T. Nees 1818, anamorphic 


1863 (H) 



(H) 

Hemicorynespora M.B. Ellis 1972, anamorphic 


1863 (H) 

Kylindria DiCosmo, S.M. Berch & W.B. 

Kendr. 1983, anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

Menispora Pers. 1822, anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

Monilochaetes Halst. ex Harter 1916, 

anamorphic Australiasca Sivan. & 

Alcorn 2002 (H)* 

Nawawia Marvanová 1980 (H) 

Paliphora Sivan. & B. Sutton 1985 (H) 

Phaeostalagmus W. Gams 1976, anamorphic 


1863 (H) 

Phialogeniculata Matsush. 1971 (H) 

Phialophora-like, anamorphic Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

Pseudobotrytis Krzemien. & Badura 1954, 

anamorphic Porosphaerella E. Müll. & 


Samuels 1982 (H) 

Pyrigemmula D. Magyar & R. Shoemaker 

2011 (H) 

Ramichloridium-like, anamorphic Chaetosphaerides 


Rattania Prabhugaonkar & Bhat 2009 (H) 

Sporidesmium Link 1809, anamorphic 

Umbrinosphaeria Réblová 1999, Miyoshiella 

Kawam. 1929 and Lecythothecium 

Réblová & Winka 2001 (H) 

Sporoschisma Berk. & Broome 1847, anamorphic 

Melanochaeta E. Müll., Harr 

& Sulmont 1969 and Chaetosphaeria 

Tul. & C. Tul. 1863 (H) 

Stachybotrys Corda 1837, anamorphic 

Melanopsamma Niessl 1876 (H) 

Thozetella Kuntze 1891 (H) 

Zanclospora S. Hughes & W.B. Kendr. 

1965, anamorphic Chaetosphaeria Tul. 

& C. Tul. 1863 (H) 

Coniochaetales Huhndorf, A.N. Mill. & F.A. 

Fernández 

Coniochaetaceae Malloch & Cain 

Cladobotryum-like, anamorphic Coniochaeta 

(Sacc.) Cooke 1887 (H) 

?Dactylaria Sacc. 1880, anamorphic Synaptospora 

Cain 1957 (H) 


Coniochaeta (Sacc.) Cooke 1887 (H) 

Phialophora-like, anamorphic Barrina 

A.W. Ramaley 1997 (H) 

Diaporthales Nannf. 

Cryphonectriaceae Gryzenh. & M.J. Wingf. 

Aurapex Gryzenh. & M.J. Wingf. 2006 

(C) 

Chrysoporthella Gryzenh. & M.J. Wingf. 

2004, anamorphic Chrysoporthe Gryzenh. 

& M.J. Wingf. 2004 (C) 

Endothiella Sacc. 1906, anamorphic Cryphonectria 

(Sacc.) Sacc. & D. Sacc. 

1905 (C) 

Foliocryphia Cheewangkoon & Crous 

2010 (C) 

Prosopidicola Crous & C.L. Lennox 2004 

(C) 

Ursicollum Gryzenh. & M.J. Wingf. 2006 

(C) 

Diaporthaceae Höhn. ex Wehm. 

Mazzantiella Höhn. 1925, anamorphic 

187

188 

Mazzantia Mont. 1855 (C) 

Phomopsis (Sacc.) Bubák 1905, anamorphic 

Diaporthe Nitschke 1870 (C) 

Gnomoniaceae G. Winter 

Asteroma DC. 1815, anamorphic Gnomoniella 

Sacc. 1881 and Plagiostoma 

Fuckel 1870 and Pleuroceras Riess. 

The type of the genus is considered to 

be Montagnellina stellaris (Pers.) 

Theiss. & Syd. 1915 (C) 

?Depazea Fr. 1818, anamorphic Pleuroceras 

Riess 1854 (C) 

Diplodina Westend. 1857, anamorphic 

Plagiostoma Fuckel 1870 (C) 

Discosporium Sacc. & P. Syd. 1902 anamorphic 

Cryptodiaporthe Petr. 1921 C. 

Tul. (C) 

Discula Sacc. 1884, anamorphic Apiognomonia 

Höhn. 1917, Gnomonia Ces. & 

De Not. 1863 and Amphiporthe Petr. 

1971 (C) 

Gloeosporidina Petr. 1921, anamorphic 

Apiognomonia Höhn. 1917 (C) 

Mazzantiella Höhn. 1925, anamorphic 

Mamianiella Höhn. 1917 (C) 

Millerburtonia Cif. 1951, anamorphic Plagiostoma 


Phaeocytostroma Petr. 1921, anamorphic 

Clypeoporthe Höhn 1919 (C) 

?Septogloeum Sacc. 1880, anamorphic 

Pleuroceras Riess 1854 (C) 

Uniseta Ciccar. 1948, anamorphic Plagiostoma 


?Zythia Fr. 1849 (C) 

Melanconidaceae G. Winter 

Dumortieria Westend. 1857 (as Cytosporina 

Sacc. 1884), anamorphic Melanamphora 

Lafl. 1976 and Melogramma 

Fr. 1849 (C) 

Harknessia Cooke 1881, anamorphic Wuestneia 

Auersw. ex Fuckel 1863 (C) 

Hendersonula Speg. 1880, anamorphic 

Pseudovalsella Höhn. 1918 and Dictyoporthe 

Petr. 1955 (C) 

Mastigosporella Höhn. 1914, anamorphic 

Wuestneiopsis J. Reid & Dowsett 1990 

(C) 

Melanconiopsis Ellis & Everh. 1900, anamorphic 

Massariovalsa Sacc. 1882 (C) 

Melanconium Link 1809, anamorphic 


Melanconis Tul. & C. Tul. 1863 (C) 

?Stilbospora Pers. 1794, anamorphic Prosthecium 

Fresen. 1852 (C) 

Tubakia B. Sutton 1973, anamorphic 

Dicarpella Syd. & P. Syd. 1921 (C) 

Pseudoplagiostomaceae Cheewangkoon, M.J. 

Wingf. & Crous 

Cryptosporiopsis-like, anamorphic Pseudoplagiostoma 

Cheewangkoon, M.J. 

Wingf. & Crous 2010 (C) 

Pseudovalsaceae M.E. Barr 

Coryneum Nees 1816, anamorphic Pseudovalsa 

Ces. & De Not. 1863 (C) 

Schizoparmeaceae Rossman 

Coniella Höhn. 1918 (C) 

Pilidiella Petr. & Syd. 1927, anamorphic 

Schizoparme Shear 1923 (C) 

Sydowiellaceae Lar. N. Vassiljeva 

Cylindrosporella Höhn. 1916, anamorphic 

Stegophora Syd. & P. Syd. 1916 (C) 

Phoma-like, anamorphic Hapalocystis 

Auersw. ex Fuckel 1863 (C) 

Togniniaceae Réblová, L. Mostert, W. Gams 

& Crous 

Phaeoacremonium W. Gams, Crous & 

M.J. Wingf. 1996, anamorphic 

Togninia Berl. 1900 (H) 

Valsaceae Tul. & C. Tul. 

Amphicytostroma Petr. 1921, anamorphic 

Amphiporthe Petr. 1971 (C) 

Cytospora Ehrenb. 1818, anamorphic 

Leucostoma (Nitschke) Höhn. 1917, 

Valsa Fr. 1849, Valsella Fuckel 1870 

and Valseutypella Höhn. 1919 (C) 

Harpostroma Höhn. 1928, anamorphic 

Leptosillia Höhn. 1928 (C) 

?Lasmenia Speg. 1886, anamorphic Chadefaudiomyces 

Kamat, Rao, Patil & 

Ullasa 1974 (C) 

Phomopsis (Sacc.) Bubák 1905, anamorphic 

Clypeoporthella Petr. 1924 (C) 

Diaporthales, genera incertae sedis 


Thailandiomyces Pinruan, 

Sakay., K.D. Hyde & E.B.G. Jones

2008 (H) 

Apoharknessia Crous & S.J. Lee 2004 (C) 

Apomelasmia Grove 1937 (C) 

Asterosporium Kunze 1819 (C) 


Thailandiomyces Pinruan, Sakay., 

K.D. Hyde & E.B.G. Jones 2008 

(H) 

Cytospora-like, anamorphic Pachytrype 

Berl. ex M.E. Barr, J.D. Rogers & Y.M. 

Ju 1993 (C) 

Dwiroopa Subram. & Muthumary 

?Greeneria Scribn. & Viala 1887 (C) 

Hormonema Lagerb. & Melin 1927, anamorphic 

Valsaria Ces. & De Not. 1863 

(H) 

Hypodermina Höhn. 1916 (C) 

Macrohilum H.J. Swart 1988 (C) 

Natarajania Pratibha & Bhat 2006 (H) 

Phomopsis-like, anamorphic Caudospora 

Starbäck 1889 (C) 

Rabenhorstia Fr. 1849, anamorphic Hercospora 

Fr. 1825 (C) 

Sirococcus Preuss 1855 (C) 

Stenocarpella Syd. & P. Syd. 1917 (C) 

Synchaetomella Decock & Seifert 2005 

(H) 

Magnaporthales Thongk., Vijakr. & K.D. 

Hyde 

Magnaporthaceae P.F. Cannon 


Clasterosphaeria Sivan. 1984 (H) 

Didymobotryum Sacc. 1886, anamorphic 

Ceratosphaerella Huhndorf, Greif, 

Mugambi & A.N. Mill. 2008 (H) 


Gaeumannomyces Arx & D.L. Olivier 

1952 and Magnaporthe R.A. Krause & 

R.K. Webster 1972 (H) 

Mycoleptodiscus Ostaz. 1968, anamorphic 

Omnidemptus P.F. Cannon & Alcorn 

1994 (H) 

Nakataea Hara 1939, anamorphic Magnaporthe 

R.A. Krause & R.K. Webster 

1972 (H) 

Pyricularia Sacc. 1880, anamorphic Magnaporthe 

R.A. Krause & R.K. Webster 

1972 (H) 

Trichocladium-like, anamorphic Gaeumannomyces 

Arx & D.L. Olivier 1952 (H) 


Ophiostomatales Benny & Kimbr. 

Kathistaceae Malloch & M. Blackw. 

Mattirolella S. Colla 1929 (C) 

Termitaria Thaxt. 1920 (H) 

Termitariopsis M. Blackw., Samson & 

Kimbr. 1980 (H) 

Ophiostomataceae Nannf. 

Acremonium-like, anamorphic Fragosphaeria 

Shear 1923 (H) 

Ambrosiella Brader ex Arx & Hennebert 

1965, anamorphic Grosmannia Goid. 

1936 and Ophiostoma Syd. & P. Syd. 

1919 (H) 

Graphium Corda 1837, anamorphic Ophiostoma 

Syd. & P. Syd. 1919 (H) 

Hyalobelemnospora Matsush. 1993, anamorphic 

Subbaromyces Hesselt. 1953 

(H) 

Hyalorhinocladiella H.P. Upadhyay & 

W.B. Kendr. 1975, anamorphic Ceratocystiopsis 

H.P. Upadhyay & W.B. 

Kendr. 1975, Klasterskya Petr. 1940 

and Ophiostoma Syd. & P. Syd. 1919 

(H) 

Hyalorhinocladiella-like, anamorphic Klasterskya 

Petr. 1940 (H) 

Leptographium Lagerb. & Melin 1927, 

anamorphic Grosmannia Gold. 1936 

(H)* 

Pesotum J.L. Crane & Schokn. 1973, 

anamorphic Ophiostoma Syd. & P. Syd. 

1919 (H) 

Pesotum-like, anamorphic Grosmannia 

Gold. 1936 (H) 

Phialographium H.P. Upadhyay & W.B. 

Kendr. 1974, anamorphic Ophiostoma 

Syd. & P. Syd. 1919 (C) 

Raffaelea Arx & Hennebert 1965, anamorphic 

Grosmannia Goid. 1936 and 

Ophiostoma Syd. & P. Syd. 1919 (H) 

Sporothrix Hektoen & C.F. Perkins 1901, 

anamorphic Ophiostoma Syd. & P. Syd. 

1919 (H) 

Sordariales Chad. ex D. Hawksw. & O.E. 

Erikss. 

Chaetomiaceae G. Winter 

Acrophialophora Edward 1961 (H) 

Botryotrichum Sacc. & Marchal 1885, anamorphic 

Chaetomium Kunze 1817 and 

Farrowia D. Hawksw. (H) 

189

190 

Humicola Traaen 1914, anamorphic Chaetomium 


Madurella Brumpt 1905 (H) 

Myceliophthora Costantin 1892, anamorphic 

Corynascus Arx 1973 and Thielavia 

Zopf 1876 (H) 

Papulaspora-like, Preuss 1851, anamorphic 

Chaetomium Kunze 1817 (H) 

Phialophora-like, anamorphic Chaetomidium 

(Zopf) Sacc. 1882 (H) 

Retroconis de Hoog & Bat. Vegte 1989 

(H) 

Taifanglania Z.Q. Liang, Y.F. Han, H.L. 

Chu & R.T.V. Fox 2009, anamorphic 

Thielavia Zopf 1876 (H) 

Trichocladium Harz 1871, anamorphic 

Chaetomium Kunze 1817 (H) 


Thielavia Zopf 1876 (H) 

Lasiosphaeriaceae Nannf. 

Angulimaya Subram. & Lodha 1964, anamorphic 

Bombardioidea C. Moreau ex 

N. Lundqv. 1972 (H) 

Bagadiella Cheewangkoon & Crous 2009 

(H) 

Cladorrhinum Sacc. & Marchal 1885, 

anamorphic Apiosordaria Arx & W. 

Gams 1967, Apodospora Cain & J.H. 

Mirza 1970, Arnium Nitschke ex G. 

Winter 1873, Cercophora Fuckel 1870 

and Podospora Ces. 1856 (H) 

Mammaria Ces. ex Rabenh. 1854, anamorphic 

Pseudocercophora Subram. & 

Sekar 1986 and Cercophora Fuckel 

1870 (H) 

Phialophora-like, anamorphic Cercophora 

Fuckel 1870, Eosphaeria Höhn. 1917, 

Lasiosphaeria Ces. & De Not. 1863 and 

Podospora Ces. 1856 (H) 

Sordariaceae G. Winter 

Chrysonilia Arx 1981, anamorphic Neurospora 

Shear & B.O. Dodge 1927 (H) 


Diplogelasinospora Cain 1961 

(H) 

Virgariella S. Hughes 1953, anamorphic 

Effetia Bartoli, Maggi & Persiani 1984 

(H) 

Sordariales, genera incertae sedis 

Kionochaeta P.M. Kirk & B. Sutton 1986 


(H) 

Lasiadelphia Réblová & W. Gams 2011 

(H) 

Lunulospora Ingold 1942 (H) 

Madurella Brumpt 1905 (H) 


Ascotaiwania Sivan. & H.S. Chang 

1992 (H) 

Monotosporella S. Hughes 1958, anamorphic 

Ascotaiwania Sivan. & H.S. 

Chang 1992 (H) 


Sordaria Ces. & De Not. 1863 (H) 

Ramophialophora M. Calduch, Stchigel, 

Gené & Guarro 2004 (H) 

Trichocladium-like, anamorphic Ascolacicola 

Ranghoo & K.D. Hyde 1998, 


1992 (H) 

Ypsilonia Lév. 1846, anamorphic Acanthotheciella 

Höhn. 1911 (H) 

Sordariomycetidae, families incertae sedis 

Amplistromataceae Huhndorf, A.N. Mill., M. 

Greif & Samuels 

Acrodontium-like, anamorphic Amplistroma 

Huhndorf, A.N. Mill., M. Greif & 

Samuels 2009 (H) 

Pseudogliomastix W. Gams 1985, anamorphic 

Wallrothiella Sacc. 1882 (H) 

Annulatascaceae S.W. Wong, K.D. Hyde & 

E.B.G. Jones 

Phaeoisaria Höhn. 1909, anamorphic 

Rhamphoria Niessl 1876 (H) 

Phialophora-like, anamorphic Teracosphaeria 

Réblová & Seifert 2007 (H) 

Cephalothecaceae Höhn. 

Acremonium-like, anamorphic Albertiniella 

Kirschst. 1936 (H) 


Cryptendoxyla Malloch & Cain 

1970 (H)* 

Phialemonium W. Gams & McGinnis 

1983 (H) 

Tritirachium Limber 1940, anamorphic 

Cephalotheca Fuckel 1871 (H) 

Helminthosphaeriaceae Samuels, Candoussau 

& Magní 

Diplococcium Grove 1885, anamorphic

Helminthosphaeria Fuckel 1870 (H) 

Endophragmiella B. Sutton 1973, anamorphic 

Echinosphaeria A.N. Mill. & 

Huhndorf 2004 (H) 

Selenosporella-like, anamorphic Ruzenia 

O. Hilber 2002 (H) 

Spadicoides S. Hughes 1958, anamorphic 

Tengiomyces Réblová 1999 (H) 

Vermiculariopsiella Bender 1932, anamorphic 

Echinosphaeria A.N. Mill. & 

Huhndorf 2004 (H) 

Sordariomycetidae, genera incertae sedis 

Sterigmatobotrys Oudem. 1886 (H)* 

Brachysporiella Bat. 1952, anamorphic 


1992 (H) 

Conioscypha Höhn. 1904, anamorphic 

Conioscyphascus Réblová & Seifert 

2004 (H) 

Endophragmiella B. Sutton 1973, anamorphic 

Phaeotrichosphaeria Sivan. 

1983 (H) 

Menisporopsis S. Hughes 1952, anamorphic 

Menisporopascus Matsush. 2003 

(H) 

Phialophora-like, anamorphic Linocarpon 

Syd. & P. Syd. 1917 (H) 

Pleurothecium Höhn. 1919, anamorphic 

Carpoligna F.A. Fernández & Huhndorf 

1999 (H) 

Ramichloridium-like, anamorphic Barbatosphaeria 

Reblova 2007 (H) 

Rhodoveronaea Arzanlou, W. Gams & 


Spadicoides S. Hughes 1958 (H) 

Sporothrix-like, anamorphic Barbatosphaeria 

Reblova 2008 (H) 

Subclass Xylariomycetidae sensu O.E. Erikss. 

& Winka 

Xylariales Nannf. 

Amphisphaeriaceae G. Winter 

Bartalinia Tassi 1900 (C) 

?Beltraniella Subram. 1952, anamorphic 

Leiosphaerella Höhn. 1919 (H) 

Bleptosporium Steyaert 1961, anamorphic 

Amphisphaeria Ces. & De Not. 1863 

(C) 

Ceratosporium Schwein. 1832, anamorphic 

Iodosphaeria Samuels, E. Müll. & 

O. Petrini 1987 (H) 


?Chaetoconis Clem. 1909, anamorphic 

Ceriospora Niessl 1876 (C) 

Discosia Lib. 1837, anamorphic Adisciso 

Kaz. Tanaka, Okane & Hosoya 2011 

(C)* 

Doliomyces Steyaert 1961 (C) 

?Hyalotiopsis Punith. 1970, anamorphic 

Ellurema Nag Raj & W.B. Kendr. 1986 

(C) 

Immersidiscosia Kaz. Tanaka, Okane & 

Hosoya 2011 (C)* 

Labridella Brenckle 1929, anamorphic 

Griphosphaerioma Höhn. 1918 (C) 

Labridium Vestergr. 1897, anamorphic 

Discostroma Clem. 1909 (C) 

Microdochium Syd. 1924, anamorphic 

Monographella Petr. 1924 (H) 

Monochaetia (Sacc.) Allesch. 1902 (C) 

Monochaetinula Muthumary, Abbas & B. 

Sutton 1986 (C) 

Monochaetiopsis L. Cai, Jeewon & K.D. 

Hyde 2003, anamorphic Dyrithiopsis L. 

Cai, R. Jeewon & K.D. Hyde 2003 (C) 

Morinia Berl. & Bres. 1889 (C) 

?Pestalotia De Not. 1841, anamorphic 

Broomella Sacc. 1883 (C) 

Pestalotiopsis Steyaert 1949, anamorphic 

Neobroomella Petr. 1947 and Pestalosphaeria 

M.E. Barr 1975 (C) 

Phlogicylindrium Crous, Summerb. & 

Summerell 2006 (H) 

Sarcostroma Cooke 1871, anamorphic 

Griphosphaerioma Höhn. 1918 (C) 

Seimatosporium Corda 1833, anamorphic 

anamorphic Discostroma Clem. 1909 

(C) 

Seiridium Nees 1816, anamorphic Blogiascospora 

Shoemaker, E. Müll. & 

Morgan-Jones 1966 and Lepteutypa 

Petr. 1923 (C) 

Selenosporella G. Arnaud ex MacGarvie 

1969, anamorphic Oxydothis Penz. & 

Sacc. 1898 and Iodosphaeria Samuels, 

E. Müll. & O. Petrini 1987 (H) 

Sporocadus Corda 1839 (C) 

Synnemapestaloides T. Handa & Y. 

Harada 2004 (H) 

?Truncatella Steyaert 1949 anamorphic 

Broomella Sacc. 1883 (C) 

Vermisporium H.J. Swart & M.A. Will. 

1983 (C) 

Zetiasplozna Nag Raj 1993 (C) 

191

192 

Diatrypaceae Nitschke 

Dumortieria Westend. 1857 (some as 

Cytosporina Sacc. 1884), anamorphic 

Cryptosphaeria Ces. & De Not. 1863, 

Diatrype Fr. 1849, Eutypa Tul. & C. 

Tul. 1863 and Eutypella (Nitschke) 

Sacc. 1875 (C) 

Libertella Desm. 1830, anamorphic 

Diatrype Fr. 1849, Diatrypella (Ces. & 

De. Not.) De. Not. 1863, Eutypa Tul. & 

C. Tul. 1863 and Eutypella (Nitschke) 

Sacc. 1875 (C) 

Diatrypaceae Nitschke 1869 

Phaeoisaria Höhn. 1909 anamorphic 

Eutypella (Nitschke) Sacc. 1875 (H) 

Hyponectriaceae Petr. 

Arthrinium Kunze 1817, anamorphic 

Physalospora Niessl (H) 

Aureobasidium Viala & G. Boyer 1891, 

anamorphic Discosphaerina Höhn. 

1917 (H) 

Beltraniella Subram. 1952, anamorphic 

Pseudomassaria Jacz. 1894 (H) 

Hormonema-like, anamorphic Discosphaerina 

Hohn 1917 (H) 

Kabatia Bubák 1904, anamorphic Discosphaerina 

Höhn. 1917 (C) 

Kabatiella Bubák 1907, anamorphic Discosphaerina 

Höhn. 1917 (H) 

Sarcophoma Höhn. 1916, anamorphic Discosphaerina 

Hohn 1917 (C) 

Selenophoma Maire 1907, anamorphic 

Discosphaerina Hohn 1917 (C) 

Graphostromataceae M.E. Barr, J.D. Rogers 

& Y.M. Ju 

Nodulisporium-like, anamorphic Graphostroma 

Piroz. 1974 (H) 

Xylariaceae Tul. & C. Tul. 

Acanthodochium Samuels, J.D. Rogers & 

Nagas., anamorphic Astrocystis Berk. & 

Broome and Collodiscula I. Hino & 

Katum. 1955 (H) 

Arthroxylaria Seifert & W. Gams 2002 

Dematophora R. Hartig 1883, anamorphic 

Rosellinia De Not. 1844 (H) 

?Dicyma Boulanger 1897, anamorphic 

Ascotricha Berk. 1838 (H) 


Geniculisynnema Okane & Nakagiri 2007, 

anamorphic Nemania Gray 1821 (H) 

Geniculosporium Chesters & Greenh. 

1964, anamorphic Entoleuca Syd. 1922, 

Leprieuria Laessøe, J.D. Rogers & 

Whalley 1989, Nemania Gray 1821, 

Phylacia Lév. 1845, Podosordaria Ellis 

& Holw. 1897, Rosellinia De Not. 1844 

and Xylaria Hill ex Schrank 1789 (H) 

Geniculosporium-like, anamorphic Halorosellinia 

Whalley, E.B.G. Jones, K.D. 

Hyde & Læssøe 2000, Jumillera J.D. 

Rogers, Y.M. Ju & San Martín 1997 

and Kretzschmaria Fr. 1849 (H) 

Hypocreodendron Henn. 1897, anamorphic 

Discoxylaria J.C. Lindq. & J.E. 

Wright 1964 (H) 

Libertella Desm. 1830 (C) 

Libertella-like, anamorphic Barrmaelia 

Rappaz 1995, Creosphaeria Theiss. 

1910, Jumillera J.D. Rogers, Y.M. Ju & 

San Martín 1997 and Lopadostoma 

(Nitschke) Traverso 1906 (C) 

Lindquistia Subram. & Chandrash. 1977, 

anamorphic Poronia Willd. 1787 (H) 

Masoniomyces J.L. Crane & Dumont 

1975, anamorphic Camillea Fr. 1849 

(H) 

Mirandina-like, anamorphic Kretzschmariella 

Viégas 1944 (H) 

Moelleroclavus Henn. 1902, anamorphic 

Xylaria Hill ex Schrank 1789 (H) 

Muscodor Worapong, Strobel & W.M. 

Hess 2001 (H) 

Nodulisporium Preuss 1849, anamorphic 

Annulohypoxylon Y.M. Ju, J.D. Rogers 

& H.M. Hsieh 2005, Biscogniauxia 

Kuntze 1891, Daldinia Ces. & De Not. 

1863, Entonaema Möller 1901, Hypoxylon 

Bull. 1791, Induratia Samuels, E. 

Müll. & O. Petrini, Pyrenomyxa Morgan 

1895 (as Pulveria Malloch & 

Rogerson 1977), Thamnomyces Ehrenb. 

1820, Theissenia Maubl. 1914, Thuemenella 

Penz. & Sacc. 1898 and 

Xylaria Hill ex Schrank (H) 

Nodulosporium-like, anamorphic Rhopalostroma 

D. Hawksw. 1977 and Ruwenzoria 

J. Fornier, M. Stadler, Læssøe & 

C. Decock 2010 (H) 

Padixonia Subram. 1972, anamorphic 

Xylaria Hill ex Schrank (H)

Rhinocladiella-like, anamorphic Obolarina 

Pouzar 1986 (H) 

Sporothrix-like, anamorphic Rostrohypoxylon 

J. Fourn. & M. Stadler 2010 and 

Ruwenzoria J. Fornier, M. Stadler, 

Læssøe & C. Decock 2010 (H) 

Triplicaria P. Karst. 1889, anamorphic 

Hypoxylon Bull. 1791 (H) 

Virgaria Nees 1816, anamorphic Ascovirgaria 

J.D. Rogers & Y.M. Ju 2002 (H) 

Virgariella-like, anamorphic Hypoxylon 

Bull. 1791 (H) 

Vivantia J.D. Rogers, Y.M. Ju & Cand. 

1996 (H) 

Xylocladium P. Syd. ex Lindau 1900, 

anamorphic Camillea Fr. 1849 (H) 

Xylocoremium J.D. Rogers 1984, anamorphic 

Xylaria Hill ex Schrank 1789 (H) 

Xylariales, genera incertae sedis 

Chaetochalara B. Sutton & Piroz. 1965 

(H) 

Chalara (Corda) Rabenh. 1844 (H)* 

Hadrotrichum Fuckel 1865 (H) 

Humicola-like, anamorphic Ascotrichella 

Valldos. & Guarro 1988 (H) 


anamorphic Phomatospora Sacc. 1875 

(H) 

Melanographium Sacc. 1913, possibly 

anamorphic Lasiobertia Sivan. 1978 

(H) 

Nodulisporium-like, anamorphic Coniolariella 

D. Garcıá, Stchigel & Guarro (H) 

Rhinocladiella-like, anamorphic Coniolariella 

D. Garcıá, Stchigel & Guarro (H) 

Selenosporella G. Arnaud ex MacGarvie 

1969, anamorphic Diatrypasimilis J. 

Zhou, & Kohlm. 2010 (H) 

Sporidesmina Subram. & Bhat 1989 (H) 

Subramaniomyces Varghese & V.G. Rao 

1980 (H) 

Sordariomycetes, orders incertae sedis 

Lulworthiales Kohlm., Spatafora & Volkm.- 

Kohlm. 

Lulworthiaceae Kohlm., Spatafora & Volkm.- 

Kohlm. 


Lindra I.M. Wilson 1956 (H) 

Cumulospora I. Schmidt 1985, anamorphic 

Lulwoana Kohlm., Volkm.- 


Kohlm., J. Campb., Spatafora & Gräfenhan 

2005 and Lulworthia G.K. 

Sutherl. 1916 (H) 

Halazoon Abdel-Aziz, Abdel-Wahab & 

Nagahama 2010, anamorphic Lulworthia 

G.K. Sutherl. 1916 (C) 

Hydea K.L. Pang & E.B.G. Jones 2010 

(H) 

Matsusporium E.B.G. Jones & K.L. Pang 

2010 (H) 

Moleospora Abdel-Wahab, Abdel-Aziz & 

Nagahama 2010 (H) 

Moromyces Abdel-Wahab, K.L. Pang, 

Nagahama, Abdel-Aziz & E.B.G. Jones 

2010 (H) 

Zalerion R.T. Moore & Meyers 1962, 

anamorphic Lulwoana Kohlm., Volkm.- 

Kohlm, J. Campb., Spatafora & Gräfenhan 

2005 (H) 

Lulworthiales, genera incertae sedis 

Halazoön Abdel-Aziz, Abdel Wahab & 

Nagah. 2011 (H) 

Phyllachorales M.E. Barr 

Phyllachoraceae Theiss. & H. Syd. 

Acerviclypeatus Hanlin 1990, anamorphic 

Ophiodothella Boyd 1934 (C) 

Baeumleria Petr. & Syd. 1927, anamorphic 

Phyllachora Nitschke ex Fuckel 

1870 (as Trabutia Sacc. & Roum. 1881) 

(C) 

Colletogloeum-like, anamorphic Coccodiella 

Hara 1909 (C) 

Cyclodomus Höhn. 1909, anamorphic 

Maculatifrons K.D. Hyde 1996 (C) 

Diachorella Höhn. 1918, anamorphic 

Diachora Müll. 1893 (C) 

Hysterodiscula Petr. 1942, anamorphic 

Orphnodactylis Malloch & A. Mallik 

1998 (C) 

Linochora Höhn. 1910, anamorphic Phyllachora 


Mycohypallage B. Sutton 1963, anamorphic 

Deshpandiella Kamat & Ullasa 

1973 (C) 

Oswaldina Rangel 1921, anamorphic Apiosphaeria 

Höhn. 1909 (C) 

Polystigmina Sacc. 1884, anamorphic Polystigma 

DC. 1815 (C) 

Pseudothiopsella Petr. 1928, anamorphic 

Pseudothiella Petr. 1928 (C) 

193

194 

Rhodosticta Woron. 1911, anamorphic 

Polystigma DC. 1815 and Stigmatula 

(Sacc.) Syd. & P. Syd. 1901 (C) 

Trichosphaeriales M.E. Barr 1983 

Trichosphaeriaceae G. Winter 1885 

Brachysporium Sacc. 1886, anamorphic 

Cryptadelphia Réblová & Seifert 2004 

(H) 

Koorchaloma Subram. 1953, anamorphic 

Kananascus Nag Raj 1984 (H) 

Stanjehughesia Subram. 1992, anamorphic 

Miyoshiella Kawam. 1929 and Umbrinosphaeria 

Réblová 1999 (H) 

Stromatographium Höhn. 1907, anamorphic 

Fluviostroma Samuels & E. Müll. 

1980 (H) 

Sordariomycetes, family incertae sedis 

Apiosporaceae K.D. Hyde, J. Fröhlich, Joanne 

E. Taylor & M.E. Barr 

Arthrinium Kunze 1817, anamorphic Apiospora 

Sacc. 1875 (H) 

Cordella Speg. 1886, anamorphic Apiospora 

Sacc. 1875 (H) 

Pteroconium Sacc. ex Grove 1914, anamorphic 

Apiospora Sacc. 1875 (H) 

Scyphospora L.A. Kantsch. 1928, anamorphic 

Apiospora Sacc. 1875 (C) 

Thyridiaceae O.E. Erikss. & J.Z. Yue 

Pleurocytospora Petr. 1923, anamorphic 

Thyridium Nitschke 1867 (C) 

Sordariomycetes, genera incertae sedis 

Cryptomycella Höhn. 1925, anamorphic 

Cryptomycina Höhn. 1977 (C) 

?Custingophora Stolk, Hennebert & Klopotek 

1968 (H) 

Didymostilbe-like, anamorphic Ornatispora 

K.D. Hyde, Goh, Joanne E. Taylor, 

J. Fröhl. 1999 (H) 

Dinemasporium Lév. 1846, anamorphic 

Phomatospora Sacc. 1875 (C) 

Ellisembia Subram. 1992 (H) 

Linkosia A. Hern. Gut. & B. Sutton 1997 

(H) 

Myrmecridium Arzanlou, W. Gams & 


Nigrospora Zimm. 1902, anamorphic 

Khuskia H.J. Huds. 1963 (H) 

Penicillifer Emden 1968, anamorphic 


Stellosetifera Matsush.1996 (H) 

Phomatosporella Tak. Kobay. & K. Sasaki 

1982, anamorphic Phomatospora Sacc. 

1875 (C) 

Ramichloridium-like, anamorphic Tectonidula 

Réblová 2009 (H) 

Sporothrix-like, anamorphic Phomatospora 

Sacc. 1875 and Tectonidula Réblová 

2009 (H) 

Subphylum PEZIZOMYCOTINA, orders 

incertae sedis 

Lahmiales O. E. Erikss. 

Lahmiaceae O. E. Erikss. 

Cyphellophora G.A. de Vries 1962, anamorphic 

Lahmia Körb. 1861 (H) 

ASCOMYCOTA, families incertae sedis 

Families and genera that cannot be 

placed in any of the classes and orders accepted 

in the present classification with a high degree 

of probability are listed below. 

Amorphothecaceae Parbery 

Hormoconis Arx & G.A. de Vries 1973, 

anamorphic Amorphotheca Parbery 

1969 (H) 

Batistiaceae Samuels & K.F. Rodrigues 

Acrostroma Seifert 1987, anamorphic 

Batistia Cif. 1958 (H) 

Pseudeurotiaceae Malloch & Cain 

Teberdinia Sogonov, W. Gams, Summerb. 

& Schroers 2005, anamorphic Pseudeurotium 

J.F.H. Beyma 1937 (H) 

Seuratiaceae Vuill. ex M.E. Barr 

Atichia Flot. 1850, anamorphic Seuratia 

Pat. 1904 (H) 

ASCOMYCOTA, genera incertae sedis 

Abropelta B. Sutton 1986 (C) 

Acarellina Bat. & H. Maia 1960 (C) 

Acaroconium Kocourk. & D. Hawksw. 

2008 (C) 

Acarocybe Syd. 1937 (H) 

Acarocybella M.B. Ellis 1960 (H) 

Acarocybellina Subram. 1992 (H) 

Acarocybiopsis J. Mena, A. Hern. Gut. & 

Mercado 1999 (H) 

Acaropeltis Petr. 1937 (C)

Achoropeltis Syd. 1929 (C) 

Aciculariella G. Arnaud 1954 (H) 

Acontium Morgan 1902 (H) 

Acremoniula G. Arnaud 1954 (H) 

Acrodictyella W.A. Baker & Partr. 2001 

(H) 

Acrodictyopsis P.M. Kirk 1983 (H) 

Acrodictys M.B. Ellis 1961 (H) 

Acrodontiella U. Braun & Scheuer 1995 

(H) 

Acrophragmis Kiffer & Reisinger 1970 

(H) 

Acrostaurus Deighton & Piroz. 1972 (H) 

Actinocladium Ehrenb. 1819 (H) 

Actinotexis Arx 1960 (C) 

Actinothecium Ces. 1854 (H) 

Actinothyrium Kunze 1823 (C) 

Acumispora Matsush. 1980 (H) 

Aegeritella Bałazy & J. Wiśn. 1974 (H) 

Aenigmatospora R.F. Castañeda, Saikawa, 

Guarro & Calduch 1999 (?H) 

Agaricodochium X.J. Liu 1981 (H) 

Agarwalomyces R.K. Verma & Kamal 

1987 (H) 

Agrabeeja Subram. 1995 (H) 

Agyriella Sacc. 1884 (H) 

Agyriellopsis Höhn. 1903 (C) 

Ahmadia Syd. 1939 (C) 

Ajrekarella Kamat & Kalani 1964 (C) 

Alatosessilispora K. Ando & Tubaki 1984 

(H) 

Alciphila Harmaja 2002 (H) 

Algonquinia R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Alloneottiosporina Nag Raj 1993 (C) 

Allothyriella Bat., Cif. & Nascim. 1959 

(C) 

Allothyrina Bat. & J.L. Bezerra 1964 (C) 

Allothyriopsis Bat., Cif. & H. Maia 1959 

(C) 

Alpakesa Subram. & K. Ramakr. 1954 (C) 

Alpakesiopsis Abbas, B. Sutton, Ghaffar & 

A. Abbas 2003 (C) 

Alveophoma Alcalde 1952 (C) 

Alysidiopsis B. Sutton 1973 (H) 

Amallospora Penz. 1897 (H) 

Amblyosporium Fresen. 1863 (H) 

Amerodiscosiella M.L. Farr 1961 (C) 

Amerodiscosiellina Bat. & Cavalc. 1966 

(C) 

Amerosporiopsis Petr. 1941 (C) 


Amerosympodula Matsush. 1996 (H) 

Amoenodochium Peláez & R.F. Castañeda 

1996 (H) 

Amoenomyces R.F. Castañeda, Saikawa & 

Hennebert 1996 (H) 

Amphichaetella Höhn. 1916 (H) 

Amphophialis R.F. Castañeda, W.B. 

Kendr. & Guarro 1998 (H) 

Amphoropycnium Bat. 1963 (C) 

Ampullicephala R.F. Castañeda, Minter & 

M. Stadler 2009 (H) 

Ampulliferina B. Sutton 1969 (H) 

Anaphysmene Bubák 1906 (C) 

Anarhyma M.H. Pei & Z.W. Yuan 1986 

(C) 

Anaselenosporella Heredia, R.F. Castañeda 

& R.M. Arias 2010 (H) 

Ancoraspora Mig. Rodr. 1982 (H) 

Ancorasporella J. Mena, Mercado & 

Heredia 1998 (H) 

Angiopomopsis Höhn. 1912 (C) 

Angulospora Sv. Nilsson 1962 (H) 

Annellodentimyces Matsush. 1985 (H) 

Annellodochium Deighton 1969 (H) 

Annellolacinia B. Sutton 1964 (C) 

Annellophora S. Hughes 1952 (H) 

Annellophorella Subram. 1962 (H) 

Annellophragmia Subram. 1963 (H) 

Annellosympodia McTaggart, R.G. Shivas 

& U. Braun 2007 (H) 

Antennopsis R. Heim 1952 (H) 

Anthopsis Fil. March., A. Fontana & Luppi 

Mosca 1977 (H) 

Anthracoderma Speg. 1888 (C) 

Antromyces Fresen. 1850 (H) 

Anulohypha Cif. 1962 (H) 

Anungitea B. Sutton 1973 (H) 

Anungitopsis R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Aoria Cif. 1962 (C) 

Aphanofalx B. Sutton 1986 (C) 

Apiocarpella Syd. & P. Syd. 1919 (C) 

Apogloeum Petr. 1954 (C) 

Aporellula B. Sutton 1986 (C) 

Aposporella Thaxt. 1920 (H) 

Arachnophora Hennebert 1963 (H) 

Arachnospora R.F. Castañeda, Minter & 

Camino 2003 (H) 

Arborillus Munt.-Cvetk. & Gómez-Bolea 

1998 (H) 

Arborispora K. Ando 1986 (H) 

Arbusculina Marvanová & Descals 1987 

195

196 

(H) 

Arcuadendron Sigler & J.W. Carmich. 

1976 (H) 

Ardhachandra Subram. & Sudha 1978 (H) 

Argopericonia B. Sutton & Pascoe 1987 

(H) 

Aristastoma Tehon 1933 (C) 

Arthrobotryum Ces. 1854 (C) 

Arthrocladium Papendorf 1969 (H) 

Arthrocristula Sigler, M.T. Dunn & J.W. 


Arthrosporium Sacc. 1880 (H) 

Arthrowallemia R.F. Castañeda, Dania 

García & Guarro 1998 (H) 

Articulophora C.J.K. Wang & B. Sutton 

1982 (H) 

Artocarpomyces Subram. 1996 (H) 

Arxiella Papendorf 1967 (H) 

Asbolisiomyces Bat. & H. Maia 1961 (C) 

Ascochytopsis Henn. 1905 (C) 

Ascochytulina Petr. 1922 (C) 

Ascospora Mont. 1849 (C) 

Ashtaangam Subram. 1995 (H) 

Aspilaima Bat. & H. Maia 1961 (C) 

Astelechia Cif. 1962 (H) 

Asterinothyriella Bat. & Cif. 1959 (C) 

Asterinothyrium Bat., Cif. & H. Maia 1959 

(C) 

Asteroconium Syd. & P. Syd. 1903 (C) 

Asteromyces F. Moreau & V. Moreau, in 

Hennenert 1962 (H) 

Asterostomopora Bat. & H. Maia 1960 (C) 

Asterostomopsis Bat., Cif. & H. Maia 1959 

(C) 

Asterostomulina Bat., J.L. Bezerra & H. 

Maia 1964 (C) 

Astronatelia Bat. & H. Maia 1962 (C) 

Atractilina Dearn. & Barthol. 1924 (H) 

Atrosetiphiale Matsush. 1995 (H) 

Aurantiosacculus Dyko & B. Sutton 1979 

(C) 

Aurosphaeria Sun J. Lee, Strobel, Eisenman, 

Geary, Vargas & S.A. Strobel 

2009 (C) 

Avesicladiella W.P. Wu, B. Sutton & 

Gange 1997 (H) 

Avettaea Petr. & Syd. 1927 (C) 

Bachmanniomyces D. Hawksw. 1981 (C) 

Bacillopeltis Bat. 1957 (C) 

Bactridiopsis Henn. 1904 (H) 

Bactridium Kunze 1817 (H) 

Bactrodesmiastrum Hol.-Jech 1984 (H) 


Bactrodesmiella M.B. Ellis 1959 (H) 

Badarisama Kunwar, J.B. Manandhar & 

J.B. Sinclair 1986 (H) 

Bahuchashaka Subram. 1978 (H) 

Bahugada K.A. Reddy & Vasant Rao 

1984 (H) 

Bahukalasa Subram. & Chandrash. 1979 

(H) 

Bahusandhika Subram. 1956 (H) 

Balaniopsis P.M. Kirk 1985 (H) 

Balanium Wallr. 1833 (H) 

Barbarosporina Ķirulis 1942 (C) 

Barnettella D. Rao & P.Rag. Rao 1964 (C) 

Basauxia Subram. 1995 (H) 

Batistina Peres 1961 (C) 

Beauveriphora Matsush 1975 (H) 

Beccopycnidium F. Stevens 1930 (C) 

Beejadwaya Subram. 1978 (H) 

Belemnospora P.M. Kirk 1981 (H) 

Bellulicauda B. Sutton 1967 (C) 

Beltrania Penz. 1882 (H) 

Beltraniomyces Monohar, D.K. Agarwal & 

Rao 2003 (H) 

Beltraniopsis Bat. & J.L. Bezerra 1960 (H) 

Beniowskia Racib. 1900 (H) 

Benjaminia S. Ahmad 1967 (C) 

Benjpalia Subram. & Bhat 1989 (H) 

Bharatheeya D'Souza & Bhat 2002 (H) 

Bhatia W.A. Baker & Morgan-Jones 2009 

(H) 

Bibanasiella R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Biflagellospora Matsush. 1975 (H) 

Biflagellosporella Matsush. 1993 (H) 

Bimeris Petr. 1949 (C) 

Bioconiosporium Bat. & J.L. Bezerra 1964 

(H) 

Biophomopsis Petr. 1931 (C) 

Bispora Corda 1837 (H) 

Bisseomyces R.F. Castañeda 1985 (H) 

Blarneya D. Hawksw., Coppins & P. 

James 1980 (H) 

Blastacervulus H.J. Swart 1988 (C) 

Blastocatena Subram. & Bhat 1989 (H) 

Blastodictys M.B. Ellis 1976 (H) 

Blastofusarioides Matsush. 1996 (H) 

Blastophorella Boedijn 1937 (H) 

Blastophorum Matsush. 1971 (H) 

Blastophragma Subram. 1995 (H) 

Blennoria Moug. & Fr. 1825 (C) 

Blennoriopsis Petr. 1919 (C) 

Blodgettia Harv. 1858 (H)

Bomplandiella Speg. 1886 (H) 

Bostrichonema Ces. 1867 (H) 

Botryoderma Papendorf & H.P. Upadhyay 

1969 (H) 

Botryodiplodina Dias & Sousa da Câmara 

1954 (C) 

Botryomonilia Goos & Piroz. 1975 (H) 

Botryosporium Corda 1831 (H) 

Brachydesmiella G. Arnaud ex S. Hughes 

1961 (H) 

Brachysporiellina Subram. & Bhat 1989 

(H) 

Brachysporiopsis Yanna, W.H. Ho & K.D. 

Hyde 2004 (H) 

Brefeldiopycnis Petr. & Cif. 1932 (H) 

Brevicatenospora R.F. Castañeda, Minter 

& Saikawa 2006 (H) 

Briosia Cavara 1888 (H) 

Brycekendrickia Nag Raj 1973 (C) 

Bulbilopycnis Matsush. 1996 (C) 

Bulbocatenospora R.F. Castañeda & Iturr. 

2000 (H) 

Bullaserpens Bat., J.L. Bezerra & Cavalc. 

1965 (C) 

Calcarispora Marvanová & Marvan 1963 

(H) 

Calcarisporiella de Hoog 1974 (H) 

Calceispora Matsush. 1975 (H) 

Callistospora Petr. 1955 (C) 

Calocline Syd. 1939 (C) 

?Calongia D. Hawksw. & Etayo 2010 (H) 

Camarographium Bubák 1916 (C) 

Camaropycnis E.K. Cash 1945 (C) 

Camarosporellum Tassi 1902 (C) 

Camarosporiopsis Abbas, B. Sutton & 

Ghaffar 2000 (C) 

Camposporidium Nawawi & Kuthub. 1988 

(H) 

Camptomeris Syd. 1927 (H) 

Campylospora Ranzoni 1953 (H) 

Candelabrum Beverw. 1951 (H) 

Candelosynnema K.D. Hyde & Seifert 

1992 (H) 

Capitorostrum Bat. 1957 (C) 

Capsicumyces Gamundí, Aramb. & 

Giaiotti 1979 (H) 

Carrismyces R.F. Castañeda & Heredia 

2000 (H) 

Castanedaea W.A. Baker & Partr. 2001 

(H) 

Catenella Bat. & Peres 1963 (C) 


Catenocuneiphora Matsush. 2003 (H) 

Catenophora Luttr. 1940 (C) 

Catenophoropsis Nag Raj & W.B. Kendr. 

1988 (C) 

Catenospegazzinia Subram. 1991 (H) 

Catenosubulispora Matsush. 1971 (H) 

Catenosynnema Kodsueb, K.D. Hyde & 

W.H. Ho 2007 (H) 

Catenulaster Bat. & C.A.A. Costa 1959 

(C) 

Catinopeltis Bat. & C.A.A. Costa 1957 

(C) 

Catinula Lév. 1848 (C) 

Ceeveesubramaniomyces Pratibha, K.D. 

Hyde & Bhat 2005 (H) 

Ceratocladium Corda 1839 (H) 

Ceratophorum Sacc. 1880 (H) 

Ceratopycnis Höhn. 1915 (C) 

Ceratosporella Höhn. 1923 (H) 

Cercosperma G. Arnaud ex B. Sutton & 

Hodges 1983 (H) 

Ceuthosira Petr. 1924 (C) 

Ceuthosporella Petr. & Syd. 1923 (C) 

Chaetendophragmia Matsush. 1971 (H) 

Chaetoblastophorum Morgan-Jones 1977 

(H) 

Chaetocytostroma Petr. 1920 (C) 

Chaetodiplis Clem. 1931 (C) 

Chaetodiplodia P. Karst. 1884 (C) 

Chaetodiplodina Speg. 1910(C) 

Chaetopeltaster Katum. 1975 (C) 

Chaetophiophoma Speg. 1910 (C) 

Chaetophoma Cooke 1878 (C) 

Chaetopsis Grev. 1825 (H) 

Chaetopyrena Pass. 1881 (C) 

Chaetoseptoria Tehon 1937 (C) 

Chalarodendron C.J.K. Wang & B. Sutton 

1984 (H) 

Chalarodes McKenzie 1991 (H) 

Chantransiopsis Thaxt. 1914 (H) 

Characonidia Bat. & Cavalc. 1965 (C) 

Charomyces Seifert 1987 (H) 

Chasakopama Manohar., Bagyan., N.K. 

Rao & Kunwar 2009 (H) 

Cheilaria Lib. 1830 (C) 

Cheiroidea W.A. Baker & Morgan-Jones 

2009 (H) 

Cheiromyceopsis Mercado & J. Mena 

1988 (H) 

Cheiromyces Berk. & M.A. Curtis 1857 

(H) 

Cheiromycina B. Sutton 1986 (H) 

197

198 

Cheiropolyschema Matsush. 1980 (H) 

Cheirospora Moug. & Fr. 1825 (C) 

Chiastospora Riess 1852 (C) 

Chikaneea B. Sutton 1973 (H) 

Chithramia Nag Raj 1988 (C) 

Chlamydomyces Bainier 1907 (H) 

Chlamydopsis Hol.-Jech. & R.F. Castañeda 

1986 (H) 

Choanatiara DiCosmo 1984 (C) 

Choreospora Constant. & R. Sant. 1987 

(H) 

Chromosporium Corda 1829 (H) 

Chrysachne Cif. 1938 (H) 

Chrysalidopsis Steyaert 1961 (C) 

Chryseidea Onofri 1981 (H) 

Ciferria Gonz. Frag. 1925 (C) 

Ciferriella Petr. 1930 (C) 

Ciferrina Petr. 1932 (C) 

Ciferriopeltis Bat. & H. Maia 1965 (C) 

Ciliochora Höhn. 1919 (C) 

Ciliochorella Syd. 1935 (C) 

Ciliophora Petr. 1929 (C) 

Ciliophorella Petr. 1940 (C) 

Ciliosporella Petr. 1927 (C) 

Circinoconis Boedijn 1942 (H) 

Circinotrichum Nees 1816 (H) 

Cirrosporium S. Hughes 1980 (C) 

Cissococcomyces Brain 1923 

Civisubramaniania Vittal & Dorai 1986 

(H) 

Cladochasiella Marvanová 1997 (H) 

Cladoconidium Bandoni & Tubaki 1985 

(H) 

Cladoniicola Diederich, van den Boom & 

Aptroot 2001 (C) 

Cladosporiella Deighton 1965 (H) 

Clasteropycnis Bat. & Cavalc. 1963 (C) 

Clathroconium Samson & H.C. Evans 

1982 (H) 

Clauzadeomyces Diederich 1994 (H) 

Clavariana Nawawi 1976 (H) 

Cleistocystis Sousa da Câmara 1931 (C) 

Cleistophoma Petr. & Syd. 1927 (C) 

Clithramia Nag Raj 1988 (C) 

Clohesyomyces K.D. Hyde 1993 (C) 

Clypeochorella Petr. 1923 (C) 

Clypeopatella Petr. 1942 (C) 

Clypeophialophora Bat. & Peres 1962 (C) 

Clypeopycnis Petr. 1925 (C) 

Clypeoseptoria F. Stevens & P.A. Young 

1925 (C) 

Clypeostagonospora Punith. 1981 (C) 


Coccogloeum Petr. 1955 (C) 

Codonmyces Calat. & Etayo 1999 (H) 

Colemaniella Agnihothr. 1974 (H) 

Coleodictyospora Charles 1929 (H) 

Coleophoma Höhn. 1907 (C) 

Coleoseptoria Petr. 1940 (C) 

Colispora Marvanová 1988 (H) 

Collembolispora Marvanová & Pascoal 

2003 (H) 

Colletoconis de Hoog & Aa 1978 (C) 

Colletosporium Link 1824 (H) 

Collostroma Petr. 1947 (C) 

Columnodomus Petr. 1941 (C) 

Columnothyrium Bubák 1916 (C) 

Comatospora Piroz. & Shoemaker 1971 

(C) 

Comocephalum Syd. 1939 (C) 

Condylospora Nawawi 1976 (H) 

Coniambigua Etayo & Diederich 1995 (C) 

Conicomyces R.C. Sinclair, Eicker & 

Morgan-Jones 1983 (H) 

Conioscyphopsis Goh & K.D. Hyde 1998 

(H) 

Coniothyrina Syd. 1912 (C) 

Coniozyma Crous 2008 (C) 

Conjunctospora Udagawa & Uchiy. 1999 

(H) 

Conostoma Bat. & J.L. Bezerra 1965 (C) 

Consetiella Hol.-Jech. & Mercado 1982 

(H) 

Coremiella Bubák & K. Krieg. 1912 (H) 

Cornucopiella Höhn. 1915 (C) 

Cornutispora Piroz. 1973 (C) 

Cornutostilbe Seifert 1990 (H) 

Corynecercospora V.K. Pal, Akhtar, N. 

Ahmad, Kamal & D.K. Agarwal 2006 

(H) 

Corynesporella Munjal & H.S. Gill 1961 

(H) 

Corynesporina Subram. 2009 (H) 

Corynesporopsis P.M. Kirk 1981 (H) 

Costanetoa Bat. & J.L. Bezerra 1963 (C) 

Craneomyces Morgan-Jones, R.C. Sinclair 

& Eicker 1987 (H) 

Creodiplodina Petr. 1957 (C) 

Creonecte Petr. 1949 (C) 

Creoseptoria Petr. 1937 (C) 

Creothyriella Bat. & C.A.A. Costa 1957 

(C) 

Cribropeltis Tehon 1933 (C) 

Crinitospora B. Sutton & Alcorn 1985 (C) 

Crucellisporiopsis Nag Raj 1983 (C)

Crustodiplodina Punith. 1988 (C) 

Cryptoceuthospora Petr. 1921 (C) 

Cryptocoryneopsis B. Sutton 1980 (H) 

Cryptocoryneum Fuckel 1870 (H) 

Cryptophialoidea Kuthub. & Nawawi 

1987 (H) 

Cryptosporium Kunze 1817 (C) 

Cryptostroma P.H. Greg. & S. Waller 

1952 (H) 

Cryptumbellata Udagawa & Uchiy. 1999 

(H) 

Ctenosporium R. Kirschner 2006 (H) 

Cubasina R.F. Castañeda 1986 (H) 

Culicidospora R.H. Petersen 1960 (H) 

Culicinomyces Couch, Romney & B. Rao 

1974 (H) 

Curucispora Matsush. 1981 (H) 

Curvulariopsis M.B. Ellis 1961 (H) 

Cyanopatella Petr. 1949 (C) 

Cyclomarsonina Petr. 1965 (C) 

Cylindrium Bonord. 1851 (H) 

Cylindrocarpostylus R. Kirschner & 

Oberw. 1999 (H) 

Cylindrodendrum Bonord. 1851 (H) 

Cylindrogloeum Petr. 1941 (C) 

Cylindromyces Manohar., D.K. Agarwal & 

N.K. Rao 2004 (H) 

Cylindrosympodium W.B. Kendr. & R.F. 

Castañeda 1990 (H) 

Cylindrothyrium Maire 1907 (C) 

Cylindroxyphium Bat. & Cif. 1963 (C) 

Cymbothyrium Petr. 1947 (C) 

Cyrenella Goch. 1981 (H) 

Cystidiella Malan 1943 (Y) 

Cystotrichiopsis B.A. Abbas, B. Sutton & 


Cytodiscula Petr. 1931 (C) 

Cytogloeum Petr. 1925 (C) 

Cytonaema Höhn. 1914 (C) 

Cytoplacosphaeria Petr. 1920 (C) 

Cytopleastrum Abbas, B. Sutton, Ghaffar 

& A. Abbas 2004 (C) 

Cytosphaera Died. 1916 (C) 

Cytosporella Sacc. 1884 (C) 

Cytostagonospora Bubák 1916 (C) 

Dactylifera Alcorn 1987 (H) 

Dactylosporium Harz 1871 (H) 

Dasysticta Speg. 1912 (C) 

Davisiella Petr. 1924 (C) 

Dearnessia Bubák 1916 (C) 

Deccanodia Singhai 1974 (C) 


Deichmannia Alstrup & D. Hawksw. 1990 

(H) 

Delortia Pat. & Gaillard 1888 (H) 

Dendroclathra Voglmayr & G. Delgado 

2001 (H) 

Dendrodomus Bubák 1915 (C) 

Dendrographiella Agnihothr. 1972 (H) 

Dendrographium Massee 1892 (H) 

Dendroseptoria Alcalde 1948 (C) 

Dendrospora Ingold 1943 (H) 

Dendrosporium Plakidas & Edgerton ex 

J.L. Crane 1972 (H) 

Dendryphiosphaera Lunghini & Rambelli 

1978 (H) 

Dennisographium Rifai 1977 (H) 

Denticularia Deighton 1972 (H) 

Dentocircinomyces R.F. Castañeda & 

W.B. Kendr. 1990 (H) 

Derexia Naras. 1970 (H) 

Descalsia A. Roldán & Honrubia 1989 (H) 

Desertella Mouch. 1979 (H) 

Desmidiospora Thaxt. 1891 (H) 

Dexhowardia J.J. Taylor 1970 (H) 

Diademospora B.E. Söderstr. & Bååth 

1979 (H) 

Dialaceniopsis Bat. 1959 (C) 

Diarimella B. Sutton 1980 (C) 

Dichelostroma Bat. & Peres 1963 (C) 

Dicholobodigitus G.P. White & Illman 

1988 (H) 

Dichotomophthora Mehrl. & Fitzp. ex 

P.N. Rao 1966 (H) 

Dichotomophthoropsis M.B. Ellis 1971 

(H) 

Dictyoarthrinium S. Hughes 1952 (H) 

Dictyocatenulata Finley & E.F. Morris 

1967 (H) 

Dictyodesmium S. Hughes 1951 (H) 

Dictyophrynella Bat. & Cavalc. 1964 (H) 

Dictyopolyschema M.B. Ellis 1976 (H) 

Dictyorostrella U. Braun 1999 (H) 

Dictyospiropes M.B. Ellis 1976 (H) 

Didymochaeta Sacc. & Ellis 1898 (C) 

Didymochaetina Bat. & J.L. Bezerra 1965 

(C) 

Didymopsis Sacc. & Marchal 1885 (H) 

Didymosporina Höhn. 1916 (C) 

Diedickea Syd. & P. Syd. 1913 (C) 

Digitatispora Doguet 1962 (H) 

Digitodochium Tubaki & Kubono 1989 

(H) 

199

200 

Digitomyces Mercado, M. Calduch & 

Gené 2003 (H) 

Digitopodium U. Braun, Heuchert & K. 

Schub. 2005 (H) 

Digitoramispora R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Dimastigosporium Faurel & Schotter 1965 

(C) 

Diplocladiella G. Arnaud ex M.B. Ellis 

1976 (H) 

Diplodinis Clem. 1931 (C) 

Diplodinula Tassi 1902 (C) 

Diplolaeviopsis Giralt & D. Hawksw. 

1991 (C) 

Diploplenodomus Died. 1912 (C) 

Diplozythiella Died. 1916 (C) 

Discomycetoidea Matsush. 1993 (H) 

Discosiellina Subram. & K.R.C. Reddy 

1972 (C) 

Discosporina Höhn. 1927 (C) 

Discotheciella Syd. & P. Syd 1917 (C) 

Discozythia Petr. 1922 (C) 

Dissitimurus E.g. Simmons, McGinnis & 

Rinaldi 1987 (H) 

Ditangifibula G.C. Adams 1996 (H) 

Dokmaia I. Promputtha 2003 (H) 

Domingoella Petr. & Cif. 1932 (H) 

Dothideodiplodia Murashk. 1927 (C) 

Dothioropsis Riedl 1974 (C) 

Drepanospora Berk. & M.A. Curtis 1875 

(H) 

Drudeola Kuntze 1891 (C) 

Drumopama Subram. 1957 (H) 

Dryosphaera Jørg. Koch & E.B.G. Jones 

1989 (C) 

Dualomyces Matsush. 1987 (H) 

Dwayabeeja Subram. 1958 (H) 

Dwayaloma Subram. 1957 (H) 

Dwayalomella Brisson, Piroz. & Pauzé 

1975 (C) 

Dwibahubeeja N. Srivast., A.K. Srivast. & 

Kamal 1995 (H) 

Dwibeeja Subram. 1995 (H) 

Dwiroopella Subram. & Muthumary 1986 

(C) 

Echinocatena R. Campb. & B. Sutton 

1977 (H) 

Echinochondrium Samson & Aa 1975 (H) 

Echinoconidiophorum Pereira-Carv. & 

Dianese 2009 (H) 

Elachopeltella Bat. & Cavalc. 1964 (C) 

Elattopycnis Bat. & Cavalc. 1964 (C) 


Elegantimyces Goh, K.M. Tsui & K.D. 

Hyde 1998 (H) 

Eleutheromycella Höhn. 1908 (C) 

Elletevera Deighton 1969 (H) 

Ellisiella Bat. 1956 (H) 

Elotespora R.F. Castañeda & Heredia 

2010 (H) 

Enantioptera Descals 1983 (H) 

Endobotrya Berk. & M.A. Curtis 1874 (C) 

Endobotryella Höhn. 1909 (C) 

Endocalyx Berk. & Broome 1876 (H) 

Endoconospora Gjaerum 1971 (H) 

Endocoryneum Petr. 1922 (C) 

Endogenospora R.F. Castañeda, O. 

Morillo & Minter 2010 (H) 

Endophragmiopsis M.B. Ellis 1966 (H) 

Endoplacodium Petr. 1949 (C) 

Endoramularia Petr. 1923 (C) 

Endosporoideus W.H. Ho, Yanna, K.D. 

Hyde & Goh 2005 (H) 

Endozythia Petr. 1959 (C) 

Enerthidium Syd. 1939 (C) 

Engelhardtiella A. Funk 1973 (H) 

Enridescalsia R.F. Castañeda & Guarro 

1998 (H) 

Enthallopycnidium F. Stevens 1925 (C) 

Entoderma Hanula, Andreadis & M. 

Blackw. 1991 (C) 

Epaphroconidia Calat. & V. Atienza 1995 

(C) 

Ephelidium C.W. Dodge & E.D. Rudolph 

1955 (C) 

Epicladonia D. Hawksw. 1981 (C) 

Epiclinium Fr. 1849 (C) 

Epicoccospora Budathoki & S.K. Singh 

1995 (?C) 

Epidermidophyton E. Lang 1879 (H) 

Epidochiopsis P. Karst. 1892 (H) 

Episporogoniella U. Braun 1994 (H) 

Epistigme Syd. 1924 (C) 

Eriocercospora Deighton 1969 (H) 

Eriocercosporella Rak. Kumar, A.N. Rai 

& Kamal ex U. Braun 1998 (H) 

Ernakulamia Subram. 1996 (H) 

Erysiphopsis Speg. 1910 (?C) 

Erythrogloeum Petr. 1953 (C) 

Esdipatilia Phadke 1981 (H) 

Esteya J.Y. Liou, J.Y. Shih & Tzean 1999 

(H) 

Eurotiopsis P. Karst. 1889 (?C) 

Evanidomus Caball. 1941 (C) 

Everhartia Sacc. & Ellis 1882 (H)

Everniicola D. Hawksw. 1982 (C) 

Eversia J.L. Crane & Schokn. 1977 (H) 

Excipularia Sacc. 1884 (H) 

Exochalara W. Gams & Hol.-Jech. 1976 

(H) 

Exophoma Weedon 1926 (C) 

Exosporella Höhn. 1912 (H) 

Exosporium Link 1809 (H) 

Falcocladium S.F. Silveira, Alfenas, Crous 

& M.J. Wingf. 1994 (H) 

Favostroma B. Sutton & E.M. Davison 

1983 (C) 

Fecundostilbum Prameela & Chowdhry 

2009 (H) 

Feltgeniomyces Dieder. 1990 (H) 

Fenestroconidia Calat. & Etayo 1999 (H) 

Filosporella Nawawi 1976 (H) 

Fissuricella Pore, D'Amatao & Ajello 

1977 (H) 

Flabellocladia Nawawi 1985 (H) 

Flabellospora Alas. 1968 (H) 

Flosculomyces B. Sutton 1978 (H) 

Frigidispora K.D. Hyde & Goh 1999 (C) 

Fujimyces Minter & Caine 1980 (C) 

Fuligomyces Morgan-Jones & Kamal 1984 

(H) 

Fumagopsis Speg. 1910 (H) 

Funicularius K.K. Baker & Zaim 1979 

(H) 

Furcaspora Bonar 1965 (C) 

Fusamen (Sacc.) P. Karst. 1890 (C) 

Fusariella Sacc. 1884 (H) 

Fusariopsis Horta 1919 (H) 

Fuscophialis B. Sutton 1977 (H) 

Fusicatena K. Matsush. & Matsush. 1996 

(H) 

Fusicolla Bonord. 1851 (H) 

Fusisporella Speg. 1910 (H) 

Fusticeps J. Webster & R.A. Davey 1980 

(H) 

Gamosporella Speg. 1888 (C) 

Gampsonema Nag Raj 1975 (C) 

Gangliophora Subram. 1992 (H) 

Gangliostilbe Subram. & Vittal 1976 (H) 

Garnaudia Borowska 1977 (H) 

Gaubaea Petr. 1942 (C) 

Geastrumia Bat. 1960 (C) 

Gelatinocrinis Matsush. 1995 (H) 

Gelatinopycnis Dyko & B. Sutton 1979 

(C) 

Geminoarcus K. Ando 1993 (H) 

Gemmulina Descals & Marvanová 1999 


(H) 

Geniopila Marvanová & Descals 1985 

(?C) 

Gerulajacta Preuss 1855 (C) 

Gilmaniella G.L. Barron 1964 (H) 

Giulia Tassi 1904 (C) 

Glaphyriopsis B. Sutton & Pascoe 1987 

(C) 

Glenosporopsis O.M. Fonseca 1943 (H) 

Glioannellodochium Matsush. 1989 (H) 

Glioblastocladium Matsush. 1989 (H) 

Globoconidiopsis G.F. Sepúlveda, Pereira- 

Carv. & Dianese 2009 (H) 

Globoconidium G.F. Sepúlveda, Pereira- 


Globuliroseum Sullia & K.R. Khan 1984 

(H) 

Gloeodes Colby 1920 (C) 

Gloeosporiella Cavara 1892 (?H) 

Gloiosphaera Höhn. 1902 (H) 

Glutinium Fr. 1849 (C) 

Glycyphila Mont. 1851(H) 

Goidanichiella G.L. Barron ex W. Gams 

2009 (H) 

Gonatobotryum Sacc. 1880 (H) 

Gonatophragmiella Rak. Kumar, A.N. Rai 

& Kamal 1994 (H) 

Gonatorrhodum Corda 1839 (H) 

Goniopila Marvanová & Descals 1985 

(H) 

Goosiella Morgan-Jones, Kamal & R.K. 

Verma 1986 (H) 

Goosiomyces N.K. Rao & Manohar. 1989 

(H) 

Gorgomyces M. Gönczöl & Révay 1985 

(H) 

Grallomyces F. Stevens 1918 (H) 

Granularia Sacc. & Ellis ex Sacc. 1882 

(H) 

Graphiothecium Fuckel 1870 (H) 

Groveolopsis Boedijn 1951 (C) 

Guedea Rambelli & Bartoli 1978 (H) 

Gymnoxyphium Cif., Bat. & I.J. Araujo 

1963 (C) 

Gyoerffyella Kol 1928 (H) 

Gyrothrix (Corda) Corda 1842 (H) 

Hadronema Syd. & P. Syd. 1909 (H) 

Hadrosporium Syd. 1938 (H) 

Halysiomyces E.G. Simmons 1981 (H) 

Halysium Corda 1837 (?H) 

Hansfordia S. Hughes 1951 (H) 

Hansfordiopeltis Bat. & C.A.A. Costa 

201

202 

1956 (C) 

Hansfordiopeltopsis M.L. Farr 1986 (C) 

Hapalosphaeria Syd. 1908 (C) 

Haplariella Syd. & P. Syd. 1908 (H) 

Haplariopsis Oudem. 1903 (H) 

Haplobasidion Erikss. 1889 (H) 

Haplolepis Syd. 1925 (C) 

Haplosporidium Speg. 1912 (C) 

Haptocara Drechsler 1975 (H) 

Harmoniella V.N. Boriss. 1981 (H) 

Harpographium Sacc. 1880 (H) 

Hawksworthiana U. Braun 1988 (H) 

Heimiodora Nicot 1960 (H) 

Helhonia B. Sutton 1980 (C) 

Helicofilia Matsush. 1983 (H) 

Helicogoosia Hol.-Jech. 1991 (H) 

Helicomina L.S. Olive 1948 (H) 

Helicominopsis Deighton 1960 (H) 

Helicorhoidion S. Hughes 1958 (H) 

Helicosingula P.S. van Wyk, Marasas, 

Baard & Knox-Dav. 1985 (H) 

Helicostilbe Höhn. 1902 (H) 

Helicothyrium I. Hino & Katum. 1961 (C) 

Helicoübisia Lunghini & Rambelli 1979 

(H) 

Heliscella Marvanová 1980 (H) 

Heliscina Marvanová 1980 (H) 

Helminthosporiomyces G.F. Sepúlveda, 

Pereira-Carv. & Dianese 2009 (H) 

Hemibeltrania Piroz. 1963 (H) 

Hemicorynesporella Subram. 1992 (H) 

Hemidothis Syd. & P. Syd. 1916 (C) 

Hemisphaeropsis Petr. 1947 (C) 

Hemisynnema Subram. 1995 (H) 

Hendersoniella Tassi 1900 (C) 

Hendersonina E.J. Butler 1913 (C) 

Hendersoniopsis Höhn. 1918 (C) 

Hendersonulina Petr. 1951 (C) 

Henicospora P.M. Kirk & B. Sutton 1980 

(H) 

Hermatomyces Speg. 1910 (H) 

Herposira Syd. 1938 (H) 

Herreromyces R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Heterocephalum Thaxt. 1903 (H) 

Heteroconidium Sawada 1944 (H) 

Heterosporiopsis Petr. 1950 (H) 

Hexacladium D.L. Olivier 1983 (H) 

Himantia Pers. 1801 (H) 

Hinoa Hara & I. Hino 1961 

Hirudinaria Ces. 1856 (H) 

Hoehneliella Bres. & Sacc. 1902 (C) 


Holubovaea Mercado 1983 (H) 

Homalopeltis Bat. & Valle 1961 (C) 

Hormiactis Preuss 1851 (H) 

Hormiscioideus M. Blackw. & Kimbr. 

1979 (H) 

Hormocephalum Syd. 1939 (H) 

Hormocladium Höhn. 1923 (H) 

Hormococcus Robak 1956 (C) 

Hormographis Guarro, Punsola & Arx 

1986 (H) 

Hughesinia J.C. Lindq. & Gamundí 1970 

(H) 

Humicolopsis Verona 1977 (H) 

Hyalocamposporium Révay & J. Gönczöl 

2007 (H) 

Hyalocladium Mustafa 1977 (H) 

Hyalocylindrophora J.L. Crane & Dumont 

1978 (H) 

Hyalodictyum Woron. 1916 (C) 

Hyalohelicomina T. Yokoy. 1974 (H) 

Hyalopleiochaeta R.F. Castañeda, Guarro 

& Cano 1996 (H) 

Hyalosynnema Matsush. 1975 (H) 

Hyalothyridium Tassi 1900 (C) 

Hyalotiastrum Abbas, B. Sutton, Ghaffar 

& Abbas 2003 (C) 

Hyalotiella Papendorf 1967 (C) 

Hyalotrochophora Finley & E.F. Morris 

1967 (H) 

Hydrometrospora J. Gönczöl & Révay 

1985 (H) 

Hymenella Fr. 1822 (H) 

Hymeniopeltis Bat. 1959 (C) 

Hymenobactron (Sacc.) Höhn. 1916 (H) 

Hymenopsis Sacc. 1886 (C) 

Hyphochlaena Cif. 1962 (H) 

Hyphodiscosia Lodha & K.R.C. Reddy 

1974 (H) 

Hyphodiscosioides Matsush. 1993 (H) 

Hyphopolynema Nag Raj 1977 (H) 

Hyphostereum Pat. 1892 (C) 

Hyphothyrium B. Sutton & Pascoe 1989 

(H) 

Hypocline Syd. 1939 (C) 

Hypogloeum Petr. 1923 (C) 

Hysteridium P. Karst. 1905 (C) 

Hysteropycnis Hilitzer 1929 (C) 

Ialomitzia Gruia 1964 (H) 

Icerymyces Brain 1923 (C) 

Idiocercus B. Sutton 1967 (C) 

Impudentia Vujanović 2003 (H) 

Inesiosporium R.F. Castañeda & W. Gams

1997 (H) 

Infundibulomyces Plaingam, Somrith. & 

E.B.G. Jones 2003 (C) 

Inifatiella R.F. Castañeda 1985 (H) 

Intercalarispora J.L. Crane & Schokn. 

1983 (H) 

Intralichen D. Hawksw. & M.S. Cole 2002 

(H) 

Ionophragmium Peres 1961 (C) 

Irpicomyces Deighton 1969 (H) 

Isariella Henn. 1908 (H) 

Ischnostroma Syd. & P. Syd. 1914 (H) 

Isthmolongispora Matsush. 1971 (H) 

Isthmophragmospora Kuthub. & Nawawi 

1992 (H) 

Isthmotricladia Matsush. 1971(H) 

Ityorhoptrum P.M. Kirk 1986 (H) 

Iyengarina Subram. 1958 (H) 

Jahniella Petr. 1921 (C) 

Janetia M.B. Ellis 1976 (H) 

Javonarxia Subram. 1995 (H) 

Jerainum Nawawi & Kuthub. 1992 (H) 

Jubispora B. Sutton & H.J. Swart 1986 

(C) 

Junctospora Minter & Hol.-Jech. 1981 (H) 

Junewangia W.A. Baker & Morgan-Jones 

2002 (H) 

Kalamarospora G. Delgado 2011 (‘2010) 

(H) 

Kalchbrenneriella Diederich & M.S. 

Christ. 2002 (H) 

Kaleidosporium Van Warmelo & B. 


Kamatella Anahosur 1969 (C) 

Kamatia V.G. Rao & Subhedar 1976 (H) 

Kameshwaromyces Kamal, R.K. Verma & 


Karsteniomyces D. Hawksw. 1980 (C) 

Kendrickiella K. Jacobs & M.J. Wingf. 

2001 (H) 

Ketubakia Kamat, Varghese & V.G. Rao 

1987 (H) 

Kiliophora Kuthub. & Nawawi 1993 (H) 

Kionocephala P.M. Kirk 1986 (H) 

Kmetia Bres. & Sacc. 1902 (H) 

Kmetiopsis Bat. & Peres 1960 (H) 

Knemiothyrium Bat. & J.L. Bezerra 1960 

(C) 

Knufia L.J. Hutchison & Unter. 1996 (H) 

Kodonospora K. Ando 1993 (H) 

Kolletes Kohlm. & Volkm.-Kohlm. 2005 

(H) 


Kontospora A. Roldán, Honrubia & Marvanová 

1990 (H) 

Koorchalomella Chona, Munjal & J.N. 

Kapoor 1958 (H) 

Korunomyces Hodges & F.A. Ferreira 

1981 (H) 

Kostermansinda Rifai 1968 (H) 

Kostermansindiopsis R.F. Castañeda 1986 

(H) 

Kramabeeja G.V. Rao & K.A. Reddy 

1981 (H) 

Kramasamuha Subram. & Vittal 1973 (H) 

Kreiseliella Braun 1991 (C) 

Krishnamyces Hosag. 2003 (C) 

Kumanasamuha P.Rag. Rao & D. Rao 

1964 (H) 

Kyphophora B. Sutton 1991 (C) 

Lacellina Sacc. 1913 (H) 

Lacellinopsis Subram. 1953 (H) 

Lachnodochium Marchal 1895 (H) 

Laciniocladium Petri 1917 (H) 

Laeviomyces D. Hawksw. 1981 (C) 

Lagenomyces Cavalc. & A.A. Silva 1972 

(C) 

Lambdasporium Matsush. 1971 (H) 

Lambinonia Sérus. & Diederich 2005 (H) 

Lamproconium (Grove) Grove 1937 (C) 

Lappodochium Matsush. 1975 (H) 

Lasiodiplodiella Zambett. 1955 (C) 

Lasiophoma Speg. 1918 (C) 

Lasiothyrium Syd. & P. Syd. 1913 (C) 

Lasmenia Speg. 1886 (C) 

Lasmeniella Petr. & Syd. 1927 (C) 

Latericonis G.V. Rao, K.A. Reddy & de 

Hoog 1984 (H) 

Lateriramulosa Matsush. 1971 (H) 

Laterispora Uecker, W.A. Ayers & P.B. 

Adams 1982 (H) 

Lauriomyces R.F. Castañeda 1990 (H) 

Lawalreea Dieder. 1990 (C) 

Leandria Rangel 1915 (H) 

Lecaniocola Brain 1923 (?C) 

Lecanostictopsis B. Sutton & Crous 1997 

(H) 

Lecophagus M.W. Dick 1990 (H) 

Lecythispora Chowdhry 1985 (?C) 


Igneocumulus A.W. Ramaley 2003 (H) 

Leeina Petr. 1923 (C) 

Leightoniomyces D. Hawksw. & B. Sutton 

1977 (H) 

Lembuncula Cif. 1954 (C) 

203

204 

Lemkea Morgan-Jones & R.C. Sinclair 

1983 (H) 

Lepisticola W. Gams 2009 (H) 

Leprieurinella Bat. & H. Maia 1961 (C) 

Leptochlamys Died. 1921(C) 

Leptodermella Höhn. 1915 (C) 

Leptodiscella Papendorf 1969 (H) 

Leptomelanconium Petr. 1923 (C) 

Leptophyllosticta I.E. Brezhnev 1939 (C) 

Leptostromella (Sacc.) Sacc. 1884 (C) 

Leptothyrella Sacc. 1885 (C) 

Leptothyrina Höhn. 1915 (C) 

Leptothyrium Kunze 1823 (C) 

Leucodochium Syd. & P. Syd. 1917 (H) 

Leucopenicillifer G.R.W. Arnold 1971 (H) 

Leuliisinea Matsush. 1985 (H) 

Libertiella Speg. & Roum. 1880 (H) 

Lichenobactridium Diederich & Etayo 

1995 (H) 

Lichenoconium Petr. & Syd. 1927 (C) 

Lichenodiplis Dyko & D. Hawksw. 1979 

(C) 

Lichenodiplisiella S.Y. Kondr. & Kudratov 

2002 (C) 

Lichenohendersonia Calat. & Etayo 2001 

(C) 

Lichenopuccinia D. Hawksw. & Hafellner 

1984 (H) 

Lichenostella Calat. & Etayo 1999 (H) 

Lichenosticta Zopf 1898 (C) 

Linochorella Syd. & P. Syd. 1912 (C) 

Listeromyces Penz. & Sacc. 1901 (H) 

Lobatopedis P.M. Kirk 1979 (H) 

Loliomyces Maire 1937 (H) 

Lomaantha Subram. 1954 (H)* 

Lomachashaka Subram. 1956 (H) 

Lunulospora Ingold 1942 (H) 

Luxuriomyces R.F. Castañeda 1988 (H) 

Luzfridiella R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Lylea Morgan-Jones 1975 (H) 

Lysotheca Cif. 1962 (C) 

Mackenziella Yanna & K.D. Hyde 2009 

(H) 

Macrodiplodia Sacc. 1884 (C) 

Macrodiplodiopsis Petr. 1922 (C) 

Macrotrichum Grev. 1825 (H) 

Mahabalella B. Sutton & S.D. Patil 1966 

(H) 

Manginella Bat. & H. Maia 1961 (C) 

Manoharachariella Bagyan., N.K. Rao & 

Kunwar 2009 (H) 


Mapletonia B. Sutton 1991 (C) 

Margarinomyces Laxa 1930 (H) 

Martinellisia V.G. Rao & Varghese 1977 

(H) 

Massalongina Bubák 1916 (C) 

Massariothea Syd. 1939 (C) 

Mastigosporium Riess 1852 (H) 

Matsushimaea Subram. 1978 (H) 

Matsushimiella R.F. Castañeda & Heredia 

2001 (H) 

Matsushimomyces V.G. Rao & Varghese 

1979 (H) 

Medusamyces G.L. Barron & Szijarto 

1990 (H) 

Megacapitula J.L. Chen & Tzean 1993 (H) 

Megalodochium Deighton 1960 (H) 

Melanocephala S. Hughes 1979 (H) 

Melanodiscus Höhn. 1918 (H) 

Melanophoma Papendorf & J.W. du Toit 

1967 (C) 

Melanosella Örösi-Pál 1936 (H) 

Melophia Sacc. 1884 (C) 

Menidochium R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Menoidea L. Mangin & Har. 1907 (H) 

Mercadomyces J. Mena 1988 (H) 

Metadiplodia Syd. 1937 (C) 

Metarhiziopsis D.W. Li, R.S. Cowles & 

Vossbrinck 2008 (H) 

Metazythia Petr. 1950 (C) 

Metazythiopsis M. Morelet 1988 (C) 

Microblastosporon Cif. 1930 (H) 

Microclava F. Stevens 1917 (H) 

Microdiscula Höhn. 1915 (C) 

Microdothiorella C.A.A. Costa & Sousa 

da Câmara 1955 (C) 

Microhendersonula Dias & Sousa da 

Câmara 1952 (C) 

Microperella Höhn. 1909 (C) 

Microtyle Speg. 1919 (C) 

Microxyphiella Speg. 1918 (C) 

Microxyphiopsis Bat. 1963 (C) 

Milospium D. Hawksw. 1975 (H) 

Mindoa Petr. 1949 (C) 

Miniancora Marvanová & Bärl. 1989 (H) 

Minimelanolocus R.F. Castañeda & Heredia 

2001 (H) 

Minimidochium B. Sutton 1969 (H) 

Minutoexcipula V. Atienza & D. Hawksw. 

1994 (H) 

Minutophoma D. Hawksw. 1981 (C) 

Mirimyces Nag Raj 1993 (C)

Mixtoconidium Etayo 1995 (C) 

Mohgaonidium Singhai 1974 (?C) 

Monochaetiella E. Castell. 1943 (C) 

Monochaetopsis Pat. 1931(H) 

Monodia Breton & Faurel 1970 (C) 

Monodidymaria U. Braun 1994 (H) 

Monodisma Alcorn 1975 (H) 

Monosporiella Speg. 1918 (H) 

Monosporium Bonord. 1851 (H) 

Monotretomyces Morgan-Jones, R.C. Sinclair 

& Eicker 1987 (H) 

Monotrichum Gäum. 1922 (H) 

Moralesia Urries 1956 (C) 

Morosporium Renault & Roche 1898 (?C) 

Morrisographium M. Morelet 1968 (H) 

Mucosetospora M. Morelet 1972 (C) 

Muiogone Thaxt. 1914 (H) 

Muirella R. Sprague 1959 (H) 

Murogenella Goos & E.F. Morris 1965 

(H) 

Mycelephas R.F. Castañeda 2009 (H) 

Mycocentrodochium K. Matsush. & Matsush. 

1996 (H) 

Mycochlamys S. Marchand & Cabral 1976 

(H) 

Mycoënterolobium Goos 1970 (H) 

Mycofalcella Marvanová, Om-Kalth. & J. 

Webster 1993 (H) 

Mycomyces Wyss-Chod. 1928 (H) 

Mycopara Bat. & J.L. Bezerra 1960 (C) 

Mycospraguea U. Braun & Rogerson 1993 

(C) 

Mycosticta Höhn. 1918 (C) 

Mycosisymbrium Carris 1994 (H) 

Mycosylva M.C. Tulloch 1973 (H) 

Mycotribulus Nag Raj & W.B. Kendr. 

1970 (C) 

Mycousteria M.L. Farr 1986 (C) 

Myiocoprula Petr. 1955 (C) 

Myriellina Höhn. 1915 (C) 

Myrmecomyces Jouvenaz & Kimbr. 1991 

(H) 

Myrotheciastrum Abbas & B. Sutton 1988 

(H) 

Mystrosporiella Munjal & Kulshr. 1969 

(H) 

Myxocephala G. Weber, Spaaij & Oberw. 

1989 (H) 

Myxoparaphysella Caball. 1941 (C) 

Myxosporella Sacc. 1881 (C) 

Myxosporidiella Negru 1960 (C) 

Myxostomellina Syd. 1931 (C) 


Myxothyriopsis Bat. & A.F. Vital 1956 (C) 

Myxothyrium Bubák & Kabát 1915 (C) 

Naemaspora Pers. 1796 (?C) 

Naemosphaera P. Karst. 1888 (C) 

Naemosphaerella Höhn. 1923 (C) 

Nagrajia R.F. Castañeda & W.B. Kendr. 

1991 (C) 

Nagrajomyces Melnik 1984 (C) 

Nakatopsis Whitton, McKenzie & K.D. 

Hyde 2001 (H) 

Nanoschema B. Sutton 1980 (C) 

Naothyrsium Bat. 1960 (C) 

Navaneethospora V.G. Rao 1994 (H) 

Necraphidium Cif. 1951 (H) 

Nematogonum Desm. 1834 (H) 

Nematographium Goid. 1935 (H) 

Nemozythiella Höhn. 1925 (C) 

Neoalpakesa Punith. 1981 (C) 

Neoarbuscula B. Sutton 1983 (H) 

Neobarclaya Sacc. 1899 (C) 

Neodiplodina Petr. 1954 (C) 

Neofuckelia Zeller & Goodd. 1935 (C) 

Neohendersonia Petr. 1921 (C) 

Neoheteroceras Nag Raj 1993 (C) 

Neojohnstonia B. Sutton 1983 (H) 

Neoligniella Naumov 1951 (C) 

Neomarssoniella U. Braun 1991 (C) 

Neomelanconium Petr. 1940 (C) 

Neoovularia U. Braun 1992 (H) 

Neopeltis Syd. 1937 (C) 

Neopericonia Kamal, A.N. Rai & Morgan- 


Neophoma Petr. & Syd. 1927 (C) 

Neoplaconema B. Sutton 1977 (C) 

Neoramularia U. Braun 1991 (H) 

Neospegazzinia Petr. & Syd. 1936 (C) 

Neosporidesmium Mercado & J. Mena 

1988 (H) 

Neottiospora Desm. 1843 (C) 

Neta Shearer & J.L. Crane 1971 (H) 

Nidulispora Nawawi & Kuthub. 1990 (H) 

Nigromacula Etayo 2002 (H) 

Nigropuncta D. Hawksw. 1981 (C) 

Nosophloea Fr. 1849 (?C) 

Nothospora Peyronel 1913 (H) 

Novozymia W.P. Wu 2005 (H) 

Nummospora E. Müll. & Shoemaker 1964 

(C) 

Nusia Subram. 1995 (H) 

Nyctalospora E.F. Morris 1972 (H) 

Obeliospora Nawawi & Kuthub. 1990 (H) 

Obstipipilus B. Sutton 1968 (C) 

205

206 

Octopodotus Kohlm. & Volkm.-Kohlm. 

2003 (C) 

Odontodictyospora Mercado 1984 (H) 

Oedothea Syd. 1930 (H) 

Ojibwaya B. Sutton 1973 (H) 

Olpitrichum G.F. Atk. 1894 (H) 

Omega B. Sutton & Minter 1988 (C) 

Oncocladium Wallr. 1833 anamorphic 

Gymnoascus Baran. 1872 (H) 

Oncopodiella G. Arnaud ex Rifai 1965 (H) 

Oncopodium Sacc. 1904 (H) 

Oncospora Kalchbr. 1880 (C) 

Oncosporella P. Karst. 1887 (C) 

Oncostroma Bat. & Marasas 1966 (C) 

Onychophora W. Gams, P.J. Fisher & J. 

Webster 1984 (H) 

Oothyrium Syd. 1939 (C) 

Ophiosira Petr. 1955 (C) 

Orbimyces Linder 1944 (H) 

Ostracodermidium Mukerji 1973 (H) 

Ovadendron Sigler & J.W. Carmich. 1976 

(H) 

Ozonium Link 1809 (H) 

Paathramaya Subram. 1956 (H) 

Pachycladina Marvanová 1987 (H) 

Pactilia Fr. 1837 (H) 

Palawaniopsis Bat., Cif. & Nascim. 1959 

Pantospora Cif. 1938 (H) 

Papilionospora V.G. Rao & B. Sutton 

1976 (H) 

Pappimyces B. Sutton & Hodges 1975 (H) 

Paraaoria R.K. Verma & Kamal 1987 

Paraarthrocladium Matsush. 1993 (H) 

Parabeltrania Rambelli 2008 (H) 

Paraceratocladium R.F. Castañeda 1987 

(H) 

Parachionomyces Thaung 1979 (H) 

Paracostantinella Subram. & Sudha 1989 

(H) 

Paracryptophiale Kuthub. & Nawawi 

1994 (H) 

Paracytospora Petr. 1925 (C) 

Paradendryphiopsis M.B. Ellis 1976 (H) 

Paradictyoarthrinium Matsush. 1996 (H) 

Paradidymobotryum C.J.K. Wang & B. 

Sutton 1984 (H) 

Paradiplodia Speg. ex Trotter 1931 (C) 

Paradischloridium Bhat & B. Sutton 1985 

(H) 

Paradiscula Petr. 1941(C) 

Paraëpicoccum Matsush. 1993 (H) 

Parafulvia Kamal, A.N. Rai & Morgan- 



Parahaplotrichum W.A. Baker & Partr. 

2001 (H) 

Paraharknessia Matsush. 2003 (H) 

Parahyalotiopsis Nag Raj 1976 (C) 

Paramassariothea Subram. & Muthumary 

1979 (C) 

Paramenisporopsis Matsush. 2003 (H) 

Parapericonia M.B. Ellis 1976 (H) 

Parapericoniella U. Braun, Heuchert & K. 

Schub. 2005 (H) 

Paraphaeoisaria de Hoog & Morgan- 


Parapithomyces Thaung 1976 (H) 

Parapleurotheciopsis P.M. Kirk 1982 (H) 

Parapyricularia M.B. Ellis 1972 (H) 

Pararobillarda Matsush. 1996 (C) 

Parasphaeropsis Petr. 1953 (C) 

Paraspora Grove 1884 (H) 

Parastigmatellina Bat. & C.A.A. Costa 

1959 (C) 

Parasympodiella Ponnappa 1975 (H) 

Paratetraploa M.K.M. Wong & K.D. 

Hyde 2002 (H) 

Paratomenticola M.B. Ellis 1976 (H) 

Paratrichoconis Deighton & Piroz. 1972 

(H) 

Paraülocladium R.F. Castañeda 1986 (H) 

Paspalomyces Linder 1933 (H) 

Patouillardiella Speg. 1889 (H) 

Patriciomyces D. Hawksw. 2001(H) 

Peethasthabeeja P.Rag. Rao 1981 (H) 

Pellionella (Sacc.) Sacc. 1902 (C) 

Peltasterinostroma Punith. 1975 (C) 

Peltasteropsis Bat. & H. Maia 1959 (C) 

Peltistroma Henn. 1904 (C) 

Peltistromella Höhn. 1907 (C) 

Peltosoma Syd. 1925 (C) 

Peltostromellina Bat. & A.F. Vital 1959 

(C) 

Peltostromopsis Bat. & A.F. Vital 1959 

(C) 

Penzigomyces Subram. 1992 (H) 

Perelegamyces R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Perizomella Syd. 1927 (C) 

Pestalozziella Sacc. & Ellis ex Sacc. 1882 

(C) 

Petrakia Syd. & P. Syd. 1913 (H) 

Petrakiopsis Subram. & K.R.C. Reddy 

1968 (H) 

Phacostroma Petr. 1955 (C)

Phacostromella Petr. 1955 (C) 

Phaeoblastophora Partr. & Morgan-Jones 

2002 (H) 

Phaeobotrys M. Calduch, Gené & Guarro 

2002 (H) 

Phaeocandelabrum R.F. Castañeda, Heredia, 

Saikawa 2009 (H) 

Phaeocytostroma Petr. 1921 (C) 

Phaeodactylella Udaiyan 1992 (H) 

Phaeodactylium Agnihothr. 1968 (H) 

Phaeodiscula Cub. 1891 (C) 

Phaeodomus Höhn. 1909 (C) 

Phaeohiratsukaea Udagawa & Iwatsu 

1990 (H) 

Phaeohymenula Petr. 1954 (H) 

Phaeoidiomyces Dorn.-Silva & Dianese 

2004 (H) 

Phaeolabrella Speg. 1912 (C) 

Phaeomonilia R.F. Castañeda, Heredia & 

R.M. Arias 2007 (H) 

Phaeomonostichella Keissl. ex Petr. 1941 

(C) 

Phaeophloeosporella Crous & B. Sutton 

1997 (C) 

Phaeophomopsis Höhn. 1917 (C) 

Phaeosphaera Bat. & Cif. 1963 (C) 

Phaeosporobolus D. Hawksw. & Hafellner 

1986 (H) 

Phaeothyrium Petr. 1947 (C) 

Phaeostilbella Höhn. 1919 (H) 

Phaeotrichoconis Subram. 1956 (H) 

Phalangispora Nawawi & J. Webster 1982 

(H) 

Phialoarthrobotryum Matsush. 1975 (H) 

Phialogangliospora Udaiyan & V.S. 

Hosag. 1992 (H) 

Phialomyces P.C. Misra & P.H.B. Talbot 

1964 (H) 

Phialophaeoisaria Matsush. 1995 (H) 

Phialoselanospora Udaiyan 1992 (H) 

Phialosporostilbe Mercado & J. Mena 

1985 (H) 

Phialostele Deighton 1969 (H) 

Phialotubus R.Y. Roy & Leelav. 1966 (H) 

Phloeoconis Fr. 1849 (H) 

Phloeosporina Höhn. 1924 (C) 

Phlyctaeniella Petr. 1922 (C) 

Phomachora Petr. & Syd. 1925 (C) 

Phomachorella Petr. 1947 (C) 

Phomyces Clem. 1931 (C) 

Phragmocephala E.W. Mason & S. 

Hughes 1951 (H) 


Phragmoconidium G.F. Sepúlveda, Pereira-Carv. 

& Dianese 2009 (H) 

Phragmogloeum Petr. 1954 (C) 

Phragmopeltis Henn. 1904 (C) 

Phragmospathula Subram. & N.G. Nair 

1966 (H) 

Phragmospathulella J. Mena & Mercado 

1986 (H) 

Phragmotrichum Kunze 1823 (C) 

Phylloedium Fr. 1825 (H) 

Phyllohendersonia Tassi 1902 (C) 

Physalidiella Rulamort 1990 (H) 

Physalidiopsis R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Pilulina G. Arnaud 1954 (H) 

Pinatubo J.B. Manandhar & Mew 1996 

(H) 

Piperivora Siboe, P.M. Kirk & P.F. Cannon 

1999 (H) 

Piricauda Bubák 1914 (H) 

Piricaudilium Hol.-Jech. 1988 (H) 

Piricaudiopsis J. Mena & Mercado 1987 

(H) 

Pirispora Faurel & Schotter 1966 (C) 

Pirostomella Sacc. 1914 (H) 

Pirozynskiella S. Hughes 2007 (H) 

Pittostroma Kowalski & T.N. Sieber 1992 

(C) 

Placella Syd. 1938 (C) 

Placodiplodia Bubák 1916 (C) 

Placonema (Sacc.) Petr. 1921 (C) 

Placonemina Petr. 1921 (C) 

Plagiostigmella Petr. 1949 (C) 

Plectonaemella Höhn. 1915 (C) 

Plectopeltis Syd. 1927 (C) 

Plectophomella Moesz 1922 (H) 

Plectophomopsis Petr. 1922 (C) 

Plectosira Petr. 1929 (C) 

Plectronidiopsis Nag Raj 1979 (C) 

Plectronidium Nag Raj 1977 (C) 

Pleiochaeta (Sacc.) S. Hughes 1951 (H) 

Plenocatenulis Bat. & Cif. 1959 (C) 

Plenophysa Syd. & P. Syd. 1920 (C) 

Plenotrichopsis Bat. 1961 (C) 

Plenotrichum Syd. 1927 (C) 

Plenozythia Syd. & P. Syd. 1916 (C) 

Pleocouturea G. Arnaud 1911 (C) 

Plesiospora Drechsler 1971 (H) 

Pleurodiscula Höhn. 1926 (C) 

Pleurodiscus Petr. 1931 (C) 

Pleurodomus Petr. 1934 (C) 

Pleuropedium Marvanová & S.H. Iqbal 

207

208 

1973 (H) 

Pleurophragmium Costantin 1888 (H) 

Pleuroplaconema Petr. 1923 (C) 

Pleuroplacosphaeria Syd. 1928 (C) 

Pleurotheciopsis B. Sutton 1973 (H) 

Pleurothyriella Petr. & Syd. 1925 (C) 

Pleurothyrium Bubák 1916 (C) 

Pleurovularia R. Kirschner & U. Braun 

2002 (H) 

Pocillopycnis Dyko & B. Sutton 1979 (C) 

Podosporiella Ellis & Everh. 1894 (H) 

Podosporium Schwein. 1832 (H) 

Poikilosperma Bat. & J.L. Bezerra 1961 

(C) 

Polyancora Voglmayr & C. Yule 2006 (H) 

Polybulbophiale Goh & K.D. Hyde 1998 

(H) 

Polycladium Ingold 1959 (H) 

Polydesmus Mont. 1845 (H) 

Polyetron Bat. & Peres 1963 (C) 

Polylobatispora Matsush. 1996 (H) 

Polyscytalum Riess 1853 (H) 

Polystomellomyces Bat. 1959 (C) 

Polystratorictus Matsush. 1993 (H) 

Polythrinciella Bat. & H. Maia 1960 (H) 

Polythrinciopsis J. Walker 1966 (H) 

Polytretophora Mercado 1983 (H) 

Porobeltraniella Gusmão 2004 (H) 

Porocladium Descals 1976 (H) 

Poropeltis Henn. 1904 (C) 

Porophilomyces U. Braun 2000 (H) 

Porosubramaniania Hol.-Jech. 1985 (H) 

Porrectotheca Matsush. 1996 (C) 

Powellia Bat. & Peres 1964 (C) 

Prathigada Subram. 1956 (H) 

Prismaria Preuss 1851 (H) 

Proboscispora Punith. 1984 (C) 

Prophytroma Sorokīn 1877 (H) 

Prosthemiella Sacc. 1881 (H) 

Protostegia Cooke 1880 (C) 

Protostegiomyces Bat. & A.F. Vital 1955 

(C) 

Protostroma Bat. 1957 (C) 

Psammina Sacc. & M. Rousseau ex E. 

Bommer & M. Rousseau 1891 (C) 

Pseudoacrodictys W.A. Baker & Morgan- 


Pseudoanguillospora S.H. Iqbal 1974 (H) 

Pseudoaristastoma Suj. Singh 1979 (C) 

Pseudoasperisporium U. Braun 2000 (H) 

Pseudobasidiospora Dyko & B. Sutton 

1978 (C) 


Pseudobeltrania Henn. 1902 (H) 

Pseudocanalisporium R.F. Castañeda & 

W.B. Kendr. 1991 (H) 

Pseudocenangium P. Karst. 1886 (C) 

Pseudochuppia Kamal, A.N. Rai & 


Pseudoclathrosphaerina Voglmayr 1997 

(H) 

Pseudoconium Petr. 1969 (C) 

Pseudocytoplacosphaeria Punith. & Spooner 

2002 (?C) 

Pseudocytospora Petr. 1923 (C) 

Pseudodichomera Höhn. 1918 (C) 

Pseudodidymaria U. Braun 1993 (H) 

Pseudodiplodia (P. Karst.) Sacc. 1884 (C) 

Pseudodiscula Laubert 1911 (C) 

Pseudoepicoccum M.B. Ellis 1971 (H) 

Pseudofuscophialis Sivan. & H.S. Chang 

1995 (H) 

Pseudogaster Höhn. 1907 (H) 

Pseudogliophragma Phadke & V.G. Rao 

1980 (H) 

Pseudographiella E.F. Morris 1966 (H) 

Pseudohendersonia Crous & M.E. Palm 

1999 (C) 

Pseudolachnea Ranoj. 1910 (C) 

Pseudoneottiospora Faurel & Schotter 

1965 (C) 

Pseudopapulaspora N.D. Sharma 1977 

(H) 

Pseudopatellina Höhn. 1908 (C) 

Pseudopeltistroma Katum. 1975 (C) 

Pseudopetrakia M.B. Ellis 1971 (H) 

Pseudophloeosporella U. Braun 1993 (C) 

Pseudophragmotrichum W.P. Wu, B. 

Sutton & Gange 1998 (C) 

Pseudopolystigmina Murashk. 1928 (C) 

Pseudoramularia Matsush. 1983 (H) 

Pseudorhizopogon Kobayasi 1983 (C) 

Pseudoschizothyra Punith. 1980 (C) 

Pseudoseptoria Speg. 1910 (C) 

Pseudosigmoidea K. Ando & N. Nakam. 

2000 (H) 

Pseudostegia Bubák 1906 (C) 

Pseudothyrium Höhn. 1927 (C) 

Pseudotorula Subram. 1958 (H) 

Pseudotracylla B. Sutton & Hodges 1976 

(C) 

Pseudotrichoconis W.A. Baker & Morgan- 


Pseudozythia Höhn. 1902 (C)

Pterulopsis Wakef. & Hansf. 1943 (H) 

Pterygosporopsis P.M. Kirk 1983 (H) 

Pucciniospora Speg. 1886 (C) 

Pulchromyces Hennebert 1973 (H) 

Pullospora Faurel & Schotter 1965 (C) 

Pulvinella A.W. Ramaley 2001 (H) 

Punctillina Toro 1934 (C) 

Pycnidioarxiella Punith. & N.D. Sharma 

1980 (C) 

Pycnidiopeltis Bat. & C.A.A. Costa 1959 

(C) 

Pycnis Bref. 1881 (C) 

Pycnodactylus Bat., A.A. Silva & Cavalc. 

1967 (C) 

Pycnodallia Kohlm. & Volkm.-Kohlm. 

2001 (?C) 

Pycnoharknessia Matsush. 1996 (C) 

Pycnomma Syd. 1924 (C) 

Pycnomoreletia Rulamort 1990 (C) 

Pycnopleiospora C.Z. Wei, Y. Harada & 

Katum. 1997 (H) 

Pycnoseynesia Kuntze 1898 (C) 

Pycnothera N.D. Sharma & G.P. Agarwal 

1974 (C) 

Pycnothyriella Bat. 1952 (C) 

Pycnothyrium Died. 1913 (C) 

Pyramidospora Sv. Nilsson 1962 (H) 

Pyrenyllium Clem. 1909 (?C) 

Pyrgostroma Petr. 1951 (C) 

Pyriculariopsis M.B. Ellis 1971 (H) 

Pyripnomyces Cavalc. 1972 (C) 

Quadracaea Lunghini, Pinzari & Zucconi 

1996 (H) 

Quadricladium Nawawi & Kuthub. 1989 

(H) 

Quasidiscus B. Sutton 1991 (C) 

Queenslandia Bat. & H. Maia 1959 (C) 

Quezelia Faurel & Schotter 1965 (C) 

Radiatispora Matsush. 1995 (H) 

Radiciseta Sawada & Katsuki 1959 (H) 

Radulidium Arzanlou, W. Gams & Crous 

2007 (H) 

Raizadenia S.L. Srivast. 1981 (H) 

Ramakrishnanella Kamat & Ullasa ex 

Ullasa 1970 (C) 

Ramicephala Voglmayr & G. Delgado 

2003 (H) 

Ramoconidiifera B. Sutton, Carmarán & 

A.I. Romero 1996 (H) 

Ranojevicia Bubák 1910 (H) 

Redbia Deighton & Piroz. 1972 (H) 

Refractohilum D. Hawksw. 1977 (H) 


Reichlingia Diederich & Scheid. 1996 (H) 

Remersonia Samson & Seifert 1997 (H) 

Repetoblastiella R.F. Castañeda, Minter & 

M. Stadler 2010 (H) 

Retiarius D.L. Olivier 1978 (H) 

Rhabdoclema Syd. 1939 (C) 

Rhabdostromella Höhn. 1915 (C) 

Rhacodiella Peyronel 1919 (H) 

Rhexoacrodictys W.A. Baker & Morgan- 


Rhexoampullifera P.M. Kirk 1982 (H) 

Rhexodenticula W.A. Baker & Morgan- 


Rhexoprolifer Matsush. 1996 (H) 

Rhinocladium Sacc. & Marchal 1885 (H) 

Rhinotrichella G. Arnaud ex de Hoog 

1977 (H) 

Rhipidocephalum Trail 1888 (H) 

Rhizohypha Chodat & Sigr. 1911 (?C) 

Rhizosphaerina B. Sutton 1986 (?C) 

Rhodesiopsis B. Sutton & R. Campb. 1979 

(C) 

Rhodothallus Bat. & Cif. 1959 (H) 

Rhombostilbella Zimm. 1902 (H) 

Rhynchodiplodia Briosi & Farneti 1906 

(C) 

Rhynchomyces Willk. 1866 (H) 

Rhynchoseptoria Unamuno 1940 (C) 

Rhynchosporina Arx 1957 (H) 

Riclaretia Peyronel 1915 (H) 

Rileya A. Funk 1979 (C) 

Robakia Petr. 1952 (C) 

Robillarda Sacc. 1882 (C) 

Rogergoosiella A. Hern. Gut. & J. Mena 

1996 (H) 

Roscoepoundia Kuntze 1898 (C) 

Rosulomyces S. Marchand & Cabral 1976 

(H) 

Rota Bat., Cif. & Nascim. 1959 (C) 

Rotaea Ces. ex Schltdl. 1851 (H) 

Rubikia H.C. Evans & Minter 1985 (C) 

Ruggieria Cif. & Montemart. 1958 (C) 

Rutola J.L. Crane & Schokn. 1978 (H) 

Sadasivania Subram. 1957 (H) 

Sanjuanomyces R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Santapauinda Subram. 1995 (H) 

Saprophragma K.B. Deshp. & K.S. Deshp. 

1966 (H) 

Saprotaphrina Verona & Rambelli 1962 

(?C) 

Sarbhoyomyces Saikia 1981 (H) 

209

210 

Sarcinodochium Höhn. 1905 (H) 

Sarcinosporon D.S. King & S.C. Jong 

1975 (H) 

Sarcopodium Ehrenb. 1818 (H) 

Satchmopsis B. Sutton & Hodges 1975 (C) 

Scaphidium Clem. 1901 (C) 

Scenomyces F. Stevens 1927 (H) 

Sceptrifera Deighton 1965 (H) 

Schizoderma Kunze 1825 (C) 

Schizothyra Bat. & C.A.A. Costa 1957 (C) 

Schizothyrella Thüm. 1880 (C) 

Schizothyropsis Bat. & A.F. Vital 1960 

(C) 

Schizotrichum McAlpine 1903 (H) 

Schroeteria G. Winter 1881 (H) 

Schwarzmannia Pisareva 1968 (C) 

Scirrhophoma Petr. 1941 (C) 

Sclerococcum Fr. 1825 (H) 

Scleroconium Syd. 1935 (H) 

Sclerodiscus Pat. 1890 (H) 

Sclerodothiorella Died. 1912 (?C) 

Sclerographiopsis Deighton 1973 (H) 

Sclerographium Berk. 1854 (H) 

Scleromeris Syd. 1926 (C) 

Scleropycnis Syd. & P. Syd. 1911 (C) 

Sclerotiella A.K. Sarbhoy & A. Sarbhoy 

1975 (H) 

Sclerozythia Petch 1937 (C) 

Scolecobasidiella M.B. Ellis 1971 (H) 

Scolecobasidium E.V. Abbott 1927 (H) 

Scolecodochium K. Matsush. & Matsush. 

1996 (H) 

Scolecotheca Søchting & B. Sutton 1997 

(C) 

Scolecozythia Curzi 1927 (C) 

Scoliotidium Bat. & Cavalc. 1963 (C) 

Scopaphoma Dearn. & House 1925 (C) 

Scopulariella Gjaerum 1971 (H) 

Scothelius Bat., J.L. Bezerra & Cavalc. 

1965 (C) 

Scutisporus K. Ando & Tubaki 1985 (H) 

Scutopeltis Bat. & H. Maia 1957 (C) 

Scutopycnis Bat. 1957 (C) 

Seifertia Partr. & Morgan-Jones 2002 (H) 

Seimatosporiopsis B. Sutton, Ghaffer & 

Abbas 1972 (C) 

Selenodriella R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Selenosira Petr. 1957 (C) 

Selenosporopsis R.F. Castañeda & W.B. 

Kendr. 1991 (H) 

Septocyta Petr. 1927 (C) 


Septocytella Syd. 1929 (C) 

Septogloeum Sacc. 1880 (C) 

Septomyxella (Höhn.) Höhn. 1923 (C) 

Septopatella Petr. 1925 (C) 

Septosporiopsis W.A. Baker & Morgan- 


Septosporium Corda 1831 (H) 

Septotrullula Höhn. 1902 (H) 

Sessiliospora D. Hawksw. 1979 (H) 

Setolibertella Punith. & Spooner 1999 

(?C) 

Setophiale Matsush. 1995 (H) 

Setosporella Mustafa & Abdul-Wahid 

1989 (H) 

Setosynnema D.E. Shaw & B. Sutton 1985 

(H) 

Seychellomyces Matsush. 1981 (H) 

Seynesiopsis Henn. 1904 (C) 

Shawiella Hansf. 1957 (C) 

Sheariella Petr. 1952 (C) 

Shrungabeeja V.G. Rao & K.A. Reddy 

1981 (H) 

Siamia V. Robert, Decock & R.F. Castañeda 

2000 (H) 

Sigmatomyces Sacc. & P. Syd. 1913 (H) 

Sirexcipula Bubák 1907 (C) 

Sirocyphis Clem. 1909 (C) 

Sirodochiella Höhn. 1925 (H) 

Sirogloea Petr. 1923 (C) 

Siroligniella Naumov 1926 (C) 

Sirophoma Höhn. 1917 (C) 

Siroplacodium Petr. 1940 (C) 

Siropleura Petr. 1934 (C) 

Siroscyphellina Petr. 1923 (C) 

Sirosperma Syd. & P. Syd. 1916 (C) 

Sirosphaera Syd. & P. Syd. 1913 (C) 

Sirosporonaemella Naumov 1951 (C) 

Sirothecium P. Karst. 1887 (C) 

Sirothyriella Höhn. 1910 (C) 

Sirothyrium Syd. & P. Syd. 1916 (C) 

Sirozythia Höhn. 1904 (C) 

Sirozythiella Höhn. 1909 (C) 

Sitochora H.B.P. Upadhyay 1964 (C) 

Slimacomyces Minter 1986 (H) 

Solicorynespora R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Soloacrospora W.B. Kendr. & R.F. Castañeda 

1991 (H) 

Solosympodiella Matsush. 1971 (H) 

Soloterminospora Matsush. 1996 (H) 

Spegazzinia Sacc. 1879 (H) 

Speiropsis Tubaki 1958 (H)

Spermatoloncha Speg. 1908 (H) 

Spermochaetella Cif. 1954 (C) 

Spermospora R. Sprague 1948 (H) 

Spermosporella Deighton 1969 (H) 

Sphaeridium Fresen. 1852 (H) 

Sphaeriostromella Bubák 1916 (C) 

Sphaeriothyrium Bubák 1916 (C) 

Sphaeromma H.B.P. Upadhyay 1964 (C) 

Sphaeronema Fr. 1815 (?C) 

Sphaerophoma Petr. 1924 (C) 

Sphaerosporium Schwein. 1832 (H) 

Sphaerulomyces Marvanová 1977 (H) 

Sphondylocephalum Stalpers 1974 (H) 

Spicularia Pers. 1822 (H) 

Spinulospora Deighton 1973 (H) 

Spiralum J.L. Mulder 1975 (H) 

Spiropes Cif. 1955 (H) 

Spondylocladiella Linder 1934 (H) 

Spondylocladiopsis M.B. Ellis 1963 (H) 

Sporidesmiopsis Subram. & Bhat 1989 (H) 

Sporocystis Morgan 1902 (H) 

Sporocephalum G. Arnaud 1952 (H) 

Sporoglena Sacc. 1894 (H) 

Sporophiala P.Rag. Rao 1970 (H) 

Sporophora Luteraan 1952 (H) 

Stachybotryella Ellis & Barthol. 1902 (H) 

Stagonopatella Petr. 1927 (C) 

Stagonopsis Sacc. 1884 (C) 

Stagonosporina Tassi 1902 (C) 

Stagonostromella Petr. & Syd. 1927 (C) 

Staheliella Emden 1974 (H) 

Stalagmochaetia Cif. & Bat. 1963 (C) 

Staphylotrichum J. Mey. & Nicot 1957 (H) 

Stauronema (Sacc.) Syd., P. Syd. & E.J. 

Butler 1916 (C) 

Stauronematopsis Abbas, B. Sutton & 


Staurophoma Höhn. 1907 (C) 

Stegolerium Strobel, W.M. Hess & E.J. 

Ford 2001 (H) 

Stegonsporiopsis Van Warmelo & B. 


Stellomyces Morgan-Jones, R.C. Sinclair 

& Eicker 1987 (H) 

Stellopeltis Bat. & A.F. Vital 1959 (C) 

Stellospora Alcorn & B. Sutton 1984 (H) 

Stellothyriella Bat. & Cif. 1959 (C) 

Stemphyliomma Sacc. & Traverso 1911 

(H) 

Stenocephalopsis Chamuris & C.J.K. 

Wang 1998 (H) 


Stenocladiella Marvanová & Descals 1987 

(H) 

Stenospora Deighton 1969 (H) 

Stephembruneria R.F. Castañeda 1988 (H) 

Stevensonula Petr. 1952 (C) 

Stichospora Petr. 1927 (C) 

Stictopatella Höhn. 1918 (C) 

Stictosepta Petr. 1964 (C) 

Stigmatellina Bat. & H. Maia 1960 (C) 

Stigmella Lév. 1842 (C) 

Stigmopeltis Syd. 1927 (C) 

Stilbellula Boedijn 1951 (H) 

Stilbophoma Petr. 1942 (C) 

Stilbospora Pers. 1794, also see Prosthecium 

(C) 

Strasseria Bres. & Sacc. 1902 (C) 

Strasseriopsis B. Sutton & Tak. Kobay. 

1970 (C) 

Stratiphoromyces Goh & K.D. Hyde 1998 

(H) 

Striosphaeropsis Verkley & Aa 1997 (C) 

Stromatopogon Zahlbr. 1897 (C) 

Stromatopycnis A.F. Vital 1956 (C) 

Stromatostysanus Höhn. 1919 (H) 

Strongylothallus Bat. & Cif. 1959 (H) 

Strumellopsis Höhn. 1909 (H) 

Stygiomyces Coppins & S.Y. Kondr. 1995 

(C) 

Stylaspergillus B. Sutton, Alcorn & P.J. 

Fisher 1982 (H) 

Subicularium M.L. Farr & Goos 1989 (H) 

Subramania D. Rao & P.Rag. Rao 1964 

(H) 

Subramanianospora Narayanan, J.K. Sharma 

& Minter 2003 (H) 

Subulispora Tubaki 1971 (H) 

Sutravarana Subram. & Chandrash. 1977 

(H) 

Suttoniella S. Ahmad 1961(C) 

Suttonina H.C. Evans 1984 (C) 

Syamithabeeja Subram. & Natarajan 1976 

(H) 

Sylviacollaea Cif. 1963 (C) 

Symbiotaphrina Kühlw. & Jurzitza ex W. 

Gams & Arx 1980 (H) 

Symphyosira Preuss 1853 (H) 

Symphysos Bat. & Cavalc. 1967 (C) 

Sympodiocladium Descals 1982 (H) 

Sympodioclathra Voglmayr 1997 (H) 

Sympodioplanus R.C. Sinclair & Boshoff 

1997 (H) 

211

212 

Synchronoblastia Uecker & F.L. Caruso 

1988 (C) 

Syncladium Rabenh. 1859 (C) 

Synnemacrodictys W.A. Baker & Morgan- 


Synnemaseimatoides K. Matsush. & 


Synnematium Speare 1920 (H) 

Synnematomyces Kobayasi 1981 (H) 

Synnemellisia N.K. Rao, Manohar. & 

Goos 1989 (H) 

Synostomina Petr. 1949 (C) 

Systremmopsis Petr. 1923 (H) 

Taeniolina M.B. Ellis 1976 (H) 

Talekpea Lunghini & Rambelli 1979 (H) 

Tandonia M.D. Mehrotra 1991 (C) 

Tandonella S.S. Prasad & R.A.B. Verma 

1970 (H) 

Tarsodisporus Bat. & A.A. Silva 1965 (C) 

Tawdiella K.B. Deshp. & K.S. Deshp. 

1966 (H) 

Tectacervulus A.W. Ramaley 1992 (C) 

Telligia Hendr. 1948 (H) 

Temerariomyces B. Sutton 1993 (H) 

Teratosperma Syd. & P. Syd. 1909 (H) 

Tetrabrachium Nawawi & Kuthub. 1987 

(H) 

Tetrabrunneospora Dyko 1978 (H) 

Tetracoccosporium Szabó 1905 (H) 

Tetranacriella Kohlm. & Volkm.-Kohlm. 

2001 (?C) 

Tetranacrium H.J. Huds. & B. Sutton 1964 

(C) 

Tetraposporium S. Hughes 1951 (H) 

Textotheca Matsush. 1996 (C) 

Thallospora L.S. Olive 1948 (H) 

Thaptospora B. Sutton & Pascoe 1987 (C) 

Tharoopama Subram. 1956 (H) 

Thirumalacharia Rathaiah 1981 (H) 

Tholomyces Matsush. 2003 (?C) 

Thoracella Oudem. 1900 (C) 

Thrinacospora Petr. 1948 (C) 

Thyriostromella Bat. & C.A.A. Costa 1959 

(C) 

Thyrostromella Höhn. 1919 (H) 

Thyrsidiella Höhn. ex Höhn. 1909 (?C) 

Thyrsidina Höhn. 1905 (C) 

Tiarosporellivora Punith. 1981 

Ticogloea G. Weber, Spaaij & W. Gams 

1994 (H) 

Tilakiopsis V.G. Rao 1994 (H) 

Titaeopsis B. Sutton & Deighton 1984 (H) 


Titaeospora Bubák 1916, also see 

Stamnaria (C) 

Tomenticola Deighton 1969 (H) 

Tompetchia Subram. 1985 (H) 

Toxosporiella B. Sutton 1986 (C) 

Toxosporiopsis B. Sutton & Sellar 1966 

(C) 

Toxosporium Vuill. 1896 (H) 

Tracylla (Sacc.) Tassi 1904 (C) 

Trematophoma Petr. 1924 (C) 

Tremellidium Petr. 1927 (C) 

Tretendophragmia Subram. 1995 (H) 

Tretocephala Subram. 1995 (H) 

Tretophragmia Subram. & Natarajan 1974 

(H) 

Tretospeira Piroz. 1972 (H) 

Tretovularia Deighton 1984 (H) 

Triadelphia Shearer & J.L. Crane 1971 

(H) 

Tribolospora D.A. Reid 1966 (C) 

Trichaegum Corda 1837 (H) 

Trichobolbus Bat. 1964 (C) 

Trichobotrys Penz. & Sacc. 1901 (H) 

Trichoconiella B.L. Jain 1976 (H) 

Trichoconis Clem. 1909 (H) 

Trichodiscula Vouaux 1910 (C) 

Trichodochium Syd. 1927 (H) 

Trichomatoclava G.F. Sepúlveda, Pereira- 


Trichomatomyces Dorn.-Silva & Dianese 

2004 (H) 

Trichomatosphaera Pereira-Carv., G.F. 

Sepúlveda & Dianese 2009 (H) 

Trichopeltulum Speg. 1889 (C) 

Trichoseptoria Cavara 1892 (C) 

Trichosporodochium Dorn.-Silva & Dianese 

2004 (H) 

Trichotheca P. Karst. 1887 (H) 

Tricladiella K. Ando & Tubaki 1984 (H) 

Tricladiopsis Descals 1982 (H) 

Tricladiospora Nawawi & Kuthub. 1988 

(H) 

Tricornispora Bonar 1967 (H) 

Trifurcospora K. Ando & Tubaki 1988 

(H) 

Triglyphium Fresen. 1852 (H) 

Trigonosporium Tassi 1900 (C) 

Tripoconidium Subram. 1978 (H) 

Triposporina Höhn. 1912 (H) 

Triramulispora Matsush. 1975 (H) 

Triscelophorus Ingold 1944 (H) 

Triscelosporium Nawawi & Kuthub. 1987

(H) 

Trisulcosporium H.J. Huds. & B. Sutton 

1964 (H) 

Troposporium Harkn. 1884 (H) 

Troposporopsis Whitton, McKenzie & 

K.D. Hyde 1999 (H) 

Trullula Ces. 1852 (C) 

Tryssglobulus B. Sutton & Pascoe 1987 

(H) 

Tuberculariopsis Höhn. 1909 (H) 

Tuberculispora Deighton & Piroz. 1972 

(H) 

Tulipispora Révay & Gönczöl 2009 (H) 

Tunicago B. Sutton & Pollack 1977 (C) 

Turturconchata J.L. Chen, T.L. Huang & 

Tzean 1999 (H) 

Tympanosporium W. Gams 1974 (H) 

Uberispora Piroz. & Hodges 1973 (H) 

Ubrizsya Negru 1965 (C) 

Ulocoryphus Michaelides, L. Hunter & 

W.B. Kendr. 1982 (H) 

Umbellidion B. Sutton & Hodges 1975 (H) 

Uncispora R.C. Sinclair & Morgan-Jones 

1979 (H) 

Urohendersonia Speg. 1902 (C) 

Urohendersoniella Petr. 1955 (C) 

Uvarispora Goos & Piroz. 1975 (H) 

Vagnia D. Hawksw. & Miądl. 1997 (C) 

Vanakripa Bhat, W.B. Kendr. & Nag Raj 

1993 (H) 

Vanbeverwijkia Agnihothr. 1961 (H) 

Vanderystiella Henn. 1908 (C) 

?Vanibandha Manohar., N.K. Rao, 

Kunwar & D.K. Agarwal 2006 (H) 

Vanterpoolia A. Funk 1982 (H) 

Variocladium Descals & Marvanová 1999 

(H) 

Vasudevella Chona, Munjal & Bajaj 1957 

(C) 

Velutipila D. Hawksw. 1987 (H) 

Ventrographium H.P. Upadhyay, Cavalc. 

& A.A. Silva 1986 (H) 

Venustocephala Matsush. 1995(H) 

Venustosynnema R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Veracruzomyces Mercado, Guarro, Heredia 

& J. Mena 2002 (H) 

Veramycella G. Delgado 2009 (H) 

Veramyces Matsush. 1993 (H) 

Verdipulvinus A.W. Ramaley 1999 (H) 

Vermispora Deighton & Piroz. 1972 (H) 

Veronaella Subram. & K.R.C. Reddy 1975 


Veronidia Negru 1964 (C) 

Verrucariella S. Ahmad 1967 (C) 

Verrucophragmia Crous, M.J. Wingf. & 

W.B. Kendr. 1994 (H) 

Verticicladus Matsush. 1993 (H) 

Vesiculohyphomyces Armando, Pereira- 


Vestigium Piroz. & Shoemaker 1972 (C) 

Viscomacula R. Sprague 1951 (H) 

Vittalia Gaws & Bhat 2007 (H) 

Vouauxiella Petr. & Syd. 1927 (C) 

Waihonghopes Yanna & K.D. Hyde 2002 

(H) 

Wardinella Bat. & Peres 1960 (C) 

Waydora B. Sutton 1976 (C) 

Websteromyces W.A. Baker & Partr. 2000 

(H) 

Weissia Bat. & M.P. Herrera 1964 (C) 

Weufia Bhat & B. Sutton 1985 (H) 

Wiesneriomyces Koord. 1907 (H) 

Wilsonomyces Adask., J.M. Ogawa & E.E. 

Butler 1990 (H) 

Wojnowicia Sacc. 1899 (C) 

Xanthoriicola D. Hawksw. 1973 (H) 

Xenidiocercus Nag Raj 1993 (C) 

Xenodomus Petr. 1922 (C) 

Xenoheteroconium Bhat, W.B. Kendr. & 

Nag Raj 1993 (H) 

Xenokylindria DiCosmo, S.M. Berch & 

W.B. Kendr. 1983 (H) 

Xenopeltis Syd. & P. Syd. 1919 (C) 

Xenoplaca Petr. 1949 (H) 

Xenostroma Höhn. 1915 (C) 

Xepicula Nag Raj 1993 (C) 

Xepiculopsis Nag Raj 1993 (C) 

Xeroconium D. Hawksw. 1981 (C) 

Xiambola Minter & Hol.-Jech. 1981 (H) 

Xiphomyces Syd. & P. Syd. 1916 (H) 

Xylochia B. Sutton 1983 (H) 

Xyloglyphis Clem. 1909 (C) 

Xylohypha (Fr.) E.W. Mason 1960 (H) 

Xylohyphopsis W.A. Baker & Partr. 2000 

(H) 

Yalomyces Nag Raj 1993 (C) 

Yinmingella Goh, K.M. Tsui & K.D. Hyde 

1999 (H) 

Ypsilina J. Webster, Descals & Marvanová 

1999 (H) 

Yuccamyces Gour, Dyko & B. Sutton 1979 

(H) 

Zakatoshia B. Sutton 1973 (H) 

Zanclospora S. Hughes & W.B. Kendr. 

213

214 

1965 (H) 

Zebrospora McKenzie 1991 (H) 

Zelandiocoela Nag Raj 1993 (C) 

Zeloasperisporium Castaneda 1996 (H) 

Zelopelta B. Sutton & R.D. Gaur 1984 (C) 

Zelosatchmopsis Nag Raj 1991 (C) 

Zelotriadelphia R.F. Castañeda, Saikawa, 

M. Stadler & Iturr. 2005 (H) 

Zetesimomyces Nag Raj 1988 (C) 

Zevadia J.C. David & D. Hawksw. 1995 

(H) 

Zilingia Petr. 1934 (C) 

Zinzipegasa Nag Raj 1993 (C) 

Zopheromyces B. Sutton & Hodges 1977 

(H) 

Zunura Nag Raj 1993 (C) 

Zygosporium Mont. 1842 (H) 

Zythia Fr. 1849 (C) 

Zyxiphora B. Sutton 1981 (H) 

Phylum BASIDIOMYCOTA R.T. Moore 

Subphylum AGARICOMYCOTINA Doweld 

Class Agaricomycetes Doweld 

Agaricales Underw. 

Agaricaceae Chevall. 

Coccobotrys Boud. & Pat. 1900, anamorphic 

Leucoagaricus Locq. ex Singer 

1948 (H) 

Fistulinaceae Lotsy 

Confistulina Stalpers 1983, anamorphic 

Fistulina Bull. 1791 (H) 

Lyophyllaceae Jülich 

Termitosphaera Cif. 1935, anamorphic 

Termitomyces R. Heim 1942 (H) 

Ugola Adans. 1763, anamorphic Asterophora 

Ditmar 1809 (H) 

Mycenaceae Overeem 

Decapitatus Redhead & Seifert 2000, 

anamorphic Mycena (Pers.) Roussel 

1806 (H) 

Niaceae Jülich 

Peyronelina P.J. Fisher, J. Webster & D.F. 

Kane 1976 (H) 

Pleurotaceae Kühner 

Antromycopsis Pat. & Trab. 1897, 

anamorphic Pleurotus (Fr.) P. Kumm. 

1871 (H) 


Nematoctonus Drechsler 1941, anamorphic 

Hohenbuehelia Schulzer 1866 (H) 

Psathyrellaceae Vilgalys, Moncalvo & 

Redhead 

Hormographiella Guarro & Gené 1992, 

Coprinellus P. Karst. 1879 (H) 

Rhacophyllus Berk. & Broome 1871, 

anamorphic Coprinopsis P. Karst. 1881 

(H) 

Tricholomataceae R. Heim ex Pouzar 

Moniliophthora H.C. Evans, Stalpers, 

Samson & Benny 1978 (H) 

Nothoclavulina Singer 1970, anamorphic 

Arthrosporella Singer 1970 (H) 

Tilachlidiopsis Keissl. 1924, anamorphic 

Dendrocollybia R.H. Petersen & Redhead 

2001 (H) 

Ugola Adans. 1763(H) 

Tilachlidiopsis Keissl. 192 (H) 

Typhulaceae Jülich 

Sclerotium Tode 1790, anamorphic 

Typhula (Pers.) Fr. 1818 (H) 

Agaricales, genera incertae sedis 

Disporotrichum Stalpers 1984 

Fibulochlamys A.I. Romero & Cabral 

1989 (H) 

Tricladiomyces Nawawi 1985 (H) 

Atheliales Jülich 

Atheliaceae Jülich 

Fibularhizoctonia G.C. Adams & Kropp 

1996, anamorphic Athelia Pers. 1822 

(H) 

Rhizoctonia DC. 1815 anamorphic Athelia 

Pers. 1822 (H) 

Sclerotium Tode 1790, anamorphic Athelia 

Pers. 1822 (H) 

Taeniospora Marvanová 1977, anamorphic 

Fibulomyces Jülich 1972 (H) 

Auriculariales J. Schröt. 1897 

Exidiaceae R.T. Moore 1978 

Ovipoculum Zhu L. Yang & R. Kirschner 

2010 (H) 

Auriculariales, genera incertae sedis 

Disporotrichum Stalpers 1984 (H) 

Oliveorhiza P. Roberts 1998, anamorphic 

Oliv-eonia Donk 1958 (H)

Cantharellales Gäum 

Botryobasidiaceae Jülich 

Haplotrichum Link 1824, anamorphic 

Botryobasidium Donk 1931 (H) 

Ceratobasidiaceae G.W. Martin 

Acanthellorhiza P. Roberts 1999, anamorphic 

Heteroacanthella 1990 (H) 

Ceratorhiza R.T. Moore 1987, anamorphic 

Ceratobasidium D.P. Rogers 1935 (H) 

Rhizoctonia DC. 1815 (and as Thanatophytum 

Nees 1816), anamorphic Helicobasidium 

Pat. 1885, Aquathanatephorus 

C.C. Tu & Kimbr. 1978 and 

Thanatephorus Donk 1956 (H) 

Hydnaceae Chevall. 

Burgella Diederich & Lawrey 2007 (H) 

Burgoa Goid. 1938, anamorphic Sistotrema 

Fr. 1821 (H) 

Ingoldiella D.E. Shaw 1972, anamorphic 

Sistotrema Fr. 1821 (H) 

Osteomorpha G. Arnaud ex Watling & 

W.B. Kendr. 1979, anamorphic Sistotrema 

Fr. 1821 and Trechispora P. 

Karst. 1890 (H) 

Oliveoniaceae P. Roberts 1998 

Oliveorhiza P. Roberts 1998, anamorphic 

Oliveonia Donk 1958 (H) 

Tulasnellaceae Juel 

Epulorhiza R.T. Moore 1987, anamorphic 

Tulasnella J. Schröt. 1888 (H) 

Cantharellales, genera incertae sedis 

Minimedusa Weresub & P.M. LeClair 

1971 (H) 

Corticiales K.H. Larss. 

Corticiaceae Herter 

Chrysorhiza T.F. Andersen & Stalpers 

1996, anamorphic Waitea Warcup & 

P.H.B. Talbot 1962 

Erythricium J. Erikss. & Hjortstam 1970 

Marchandiomyces Dieder. & D. Hawksw. 

1990, anamorphic Marchandiobasidium 

Diederich & Schultheis 2003 (H) 

Michenera Berk. & M.A. Curtis 1868, 

anamorphic Licrostroma P.A. Lemke 

1964 (H) 

Tretopileus B.O. Dodge 1946 (H) 


Hymenochaetales Oberw. 

Schizoporaceae Jülich 

Echinodia Pat. 1918, anamorphic Echinoporia 

Ryvarden 1980 (H) 

Hymenochaetales, genera incertae sedis 

Caeruleomyces Stalpers 2000 

Polyporales Gäum. 

Corticiaceae Herter 

Necator Massee 1898 (H) 

Fomitopsidaceae Jülich 

Ptychogaster Corda 1838, anamorphic 

Oligoporus Bref. 1888 and Postia Fr. 

1874 (H) 

Sporotrichum Link 1809, anamorphic 

Laetiporus Murrill 1904 and Pycnoporellus 

Murrill 1905 (H) 

Ganodermataceae Donk 

Thermophymatospora Udagawa, Awao & 

Abdullah 1986, anamorphic Ganoderma 

P. Karst. 1881 (H) 

Hyphodermataceae Jülich 

Aegerita Pers. 1801, anamorphic Bulbillomyces 

Jülich 1974 (H) 

Aegeritina Jülich 1984, anamorphic 

Subulicystidium Parmasto 1968 (H) 

Meruliaceae P. Karst. 

Bornetina L. Mangin & Viala 1903, 

anamorphic Diacanthodes Singer 1945 

(H) 

Sporotrichopsis Staplers (2000), anamorphic 

Abortiporus Murrill (H) 

Phanerochaetaceae Jülich 

Erythricium J. Erikss. & Hjortstam 1970, 

anamorph of Phanerochaete P. Karst. 

1889 

Sporotrichum Link 1809, anamorphic 

Phanerochaete P. Karst. 1889 (H) 

Polyporaceae Fr. ex Corda 

Digitellus Paulet 1791, anamorphic Lentinus 

Fr. 1825 (H) 

Mycelithe Gasp. 1841, anamorphic Polyporus 

P. Micheli ex Adans. 1763 (C)? 

Pachyma Fr. 1822, anamorphic Lentinus 

Fr. 1825, Macrohyporia I. Johans. & 

215

216 

Ryvarden 1979, Polyporus P. Micheli 

ex Adans. 1763, Lignosus Lloyd ex 

Torrend 1920 and Wolfiporia Ryvarden 

& Gilb. 1984 (H) 

Russulales Kreisel ex P.M. Kirk, P.F. Cannon 

& J.C. David 

Bondarzewiaceae Kotl. & Pouzar 

Spiniger Stalpers 1974, anamorphic 

Bondarzewia Singer 1940 and Heterobasidion 

Bref. 1888 (H) 

Echinodontiaceae Donk 

Spiniger Stalpers 1974, anamorphic Laurilia 

Pouzar 1959 (H) 

Lachnocladiaceae D.A. Reid 

Spiniger Stalpers 1974, anamorphic 

Dichostereum Pilát 1926 (H) 

Stereaceae Pilát 

Acaromyces Boekhout, Scorzetti, Gerson 

& Sztejnb. 2003 (H) 

Aleurocystis Lloyd ex G. Cunn. 1956, 

anamorphic Licrostroma P.A. Lemke 

1964 (?C) 

Matula Massee 1888, anamorphic Aleurocystis 

Lloyd ex G. Cunn.1956 (H) 

Sebacinales M. Weiss, Selosse, Rexer, A. Urb. 

& Oberw. 

Sebacinaceae K. Wells & Oberw. 

Craterocolla Bref. 1888 

Ditangium P. Karst. 1867, anamorphic 

Craterocolla Bref. 1888 (H) 

?Chaetospermum Sacc. 1892, anamorphic 

Efibulobasidium K. Wells 1975 (C) 

Flahaultiella Seifert 2009, anamorphic 

Sebacina Tul. & C. Tul. 1871 (H) 

Opadorhiza T.F. Andersen & R.T. Moore 

1996, anamorphic Sebacina Tul. & C. 

Tul. 1871 (H) 

Piriformospora Sav. Verma, Aj. Varma, 

Rexer, G. Kost & P. Franken 1998 (H) 

Thelephorales Corner ex Oberw. 

Thelephoraceae Chevall. 

Parahaplotrichum W.A. Baker & Partr. 

2001, anamorphic Botryobasidium Rick 

1959 (H) 

Agaricomycetes, genera incertae sedis 


Akenomyces G. Arnaud ex D. Hornby 

1984 (H) 

Arthrodochium R.F. Castañeda & W.B. 

Kendr. 1990 (H) 

Arualis Katz 1980 (H) 

Bartheletia G. Arnaud ex Scheuer, R. 

Bauer, M. Lutz, Stabenth., Melnik & 

Grube 2008 (H) 

Cenangiomyces Dyko & B. Sutton 1979 

(H) 

Cruciger R. Kirschner & Oberw. 1999 (H) 

Dendrosporomyces Nawawi, J. Webster & 

R.A. Davey 1977 (H) 

Ellula Nag Raj 1980 (C) 

Fibulocoela Nag Raj 1978 (C) 

Fibulotaeniella Marvanová & Bärl. 1988 

(H) 

Geotrichopsis Tzean & Estey 1991 (H) 

Gloeosynnema Seifert & G. Okada 1988 

(H) 

Glomerulomyces A.I. Romero & S.E. 

López 1989 (H) 

Glutinoagger Sivan. & Watling 1980 (H) 

Myriococcum Fr. 1823 (H) 

Nyctalina G. Arnaud 1952 (H) 

Pagidospora Drechsler 1960 (H) 

Pycnovellomyces R.F. Castañeda 1987 (C) 

Riessia Fresen. 1852 (H) 

Riessiella Jülich 1985 (H) 

Titaeella G. Arnaud ex K. Ando & Tubaki 

1985 (H) 


Class Dacrymycetes Doweld 

Dacrymycetales Henn. 

Dacrymycetaceae J. Schröt. 

Cerinosterus R.T. Moore 1987 (H) 

Dacryoscyphus R. Kirschner & Zhu L. 

Yang 2005 (H) 

Class Tremellomycetes Hibbett, Matheny, & 

Manfr. Binder 

Cystofilobasidiales Fell, Roeijmans & 

Boekhout 

Cystofilobasidiaceae K. Wells & Bandoni 

Guehomyces Fell & Scorzetti 2004 

Itersonilia Derx 1948 

Mrakiella Margesin & Fell 2008 (H) 

Phaffia M.W. Mill., Yoney. & Soneda 

1976, anamorphic Xanthophyllomyces 

Golubev 1995 (C)

Rhodozyma Phaff, M.W. Mill., Yoney. & 

Soneda 1972, anamorphic Xanthophyllomyces 

Golubev 1995 

Tausonia Babeva 1998 

Udeniomyces Nakase & Takem. 1992 (Y) 

Filobasidiales Jülich 

Filobasidiaceae L.S. Olive 

Cryptococcus Vuill. 1901, anamorphic 

Filobasidium L.S. Olive 1968 (Y) 

Tremellales Fr. 

Cuniculitremaceae J.P. Samp., R. Kirschner & 

M. Weiss 

Fellomyces Y. Yamada & I. Banno 1984 

(Y) 

Kockovaella Nakase, I. Banno & Y. 

Yamada 1991 (Y) 

Sterigmatosporidium G. Kraep. & U. 

Schulze 1983, anamorphic Cuniculitrema 

J.P. Samp. & R. Kirschner 2001 

(Y) 

Tremellina Bandoni 1986, anamorphic 

Cuniculitrema J.P. Samp. & R. 

Kirschner 2001 (H) 

Hyaloriaceae Lindau 

Helicomyxa R. Kirschner & Chee J. Chen 

2004 (H) 

Tremellaceae Fr. 

Anastomyces W.P. Wu, B. Sutton & 

Gange 1997 (H) 

Bullera Derx 1930 (Yeast) 

Cryptococcus Vuill. 1901, anamorphic 

Filobasidiella Kwon-Chung 1976 (Y) 

Hormomyces Bonord. 1851, anamorphic 

Tremella Pers. 1794 (H) 

Hormomyces-like anamorphic Biatoropsis 

Räsänen 1934 (H) 

Tsuchiyaea Y. Yamada, H. Kawas., Itoh, I. 

Banno & Nakase 1988 (H) 

Trichosporonaceae Nann. 

Asterotremella Prillinger, Lopandic & 

Sugita 2007 (Y) 

Trichosporon Behrend 1890 (Y) 

Tritirachium Limber 1940, anamorphic 

Neorehmia Höhn. 1902 (H) 

Tremellales genera incertae cedis 


Derxomyces F.Y. Bai & Q.M. Wang 2008 

(Y) 

Hannaella F.Y. Bai & Q.M. Wang 2008 

(Y) 

Moniliella Stolk & Dakin 1966 (H) 

Tremellomycetes, genera incertae sedis 

Heteromycophaga P. Roberts 1997 (Yeastlike) 

Moniliella Stolk & Dakin 1966 (H) 

Trichosporonoides Haskins & J.F.T. 

Spencer 1967 (H) 

AGARICOMYCOTINA, genera incertae sedis 

Microstella K. Ando & Tubaki 1984 (H) 

Subphylum PUCCINIOMYCOTINA R. 

Bauer, Begerow, J.P. Samp., M. Weiss & 

Oberw 

Class Agaricostilbomycetes R. Bauer, Begerow, 

J.P. Samp., M. Weiss & Oberw. 

Agaricostilbales Oberw. & R. Bauer 

Agaricostilbaceae Oberw. & R. Bauer 

Agaricostilbum J. Wright 1970 (H) 

Bensingtonia Ingold 1986 (Y) 

Sterigmatomyces Fell 1966 (Y) 

Chionosphaeraceae Oberw. & Bandoni 

Kurtzmanomyces Y. Yamada, Itoh, H. 

Kawas., I. Banno & Nakase 1989 (Y) 

Mycogloea L.S.Olive 1950 (Y) 

Class Atractiellomycetes R. Bauer, Begerow, 


Atractiellales Oberw. & Bandoni 

Phleogenaceae Gäum. 

Atractiella Sacc. 1886 (H) 

Basidiopycnides J. Reid, Eyjólfsd. & 

Georg Hausner 2008, anamorphic Basidiopycnis 

Oberw., R. Kirschner, 

R.Bauer, Begerow & Arenal 2006 (H) 

Saccoblastiaceae Jülich 

Infundibura Nag Raj & W.B. Kendr. 1981 

(H) 

Atractiellales, genera incertae sedis 

Atractogloea Oberw. & Bandoni 1982 (H) 

Hobsonia Berk. ex Massee 1891 (H) 

Leucogloea R. Kirschner 2004 (H) 


217

Pachnocybales R. Bauer, Begerow, J.P. Samp., 

M. Weiss & Oberw. 2006 

Pachnocybaceae Oberw. & R. Bauer 1989 

Pachnocybe Berk. 1836 (H) 

Class Classiculomycetes R. Bauer, Begerow, 


Classiculales R. Bauer, Begerow, Oberw. & 

Marvanová 

Classiculaceae Bauer, Begerow, Oberw. & 

Marvanová 

Jaculispora H.J. Huds. & Ingold 1960 (H) 

Naiadella Marvanová & Bandoni 1987, 

anamorphic Classicula R. Bauer, Begerow, 

Oberw. & Marvanová 2003 (H) 

Class Cryptomycocolacomycetes R. Bauer, 

Begerow, J.P. Samp., M. Weiss & Oberw. 

Cryptomycocolacales Oberw. & R. Bauer 

Cryptomycocolacaceae Oberw. & R. Bauer 

Colacosiphon R. Kirschner, R. Bauer & 

Oberw. 2001 (H) 

Class Cystobasidiomycetes R. Bauer, 

Begerow, J.P. Samp., M. Weiss & Oberw. 

Erythrobasidiales, genera incertae sedis 

Bannoa Hamam. 2002 (Y) 

Erythrobasidium Hamam., Sugiy. & Komag. 

1988 (Y) 

Class Microbotryomycetes R. Bauer, Begerow, 


Heterogastridiales Oberw. & R. Bauer 

Heterogastridiaceae Oberw. & R. Bauer 

Hyalopycnis Höhn. 1918, anamorphic 

Heterogastridium Oberw. & R. Bauer 

1990 (C) 

Leucosporidiales J.P. Samp., M. Weiss & R. 

Bauer 

Leucosporidiaceae Jülich 

Leucosporidiella Samp. 2003 (Y) 

Microbotryales R. Bauer & Oberw., genera 

incertae sedis 

Reniforma Pore & Sorenson 1990 (Y) 

Sporidiobolales Doweld 

Sporidiobolaceae R.T. Moore 

Blastoderma B. Fisch. & Brebeck 1894, 

anamorphic Sporidiobolus Nyland 1950 

(Y) 

218 


Rhodomyces Wettst. 1885, anamorphic 

Sporidiobolus Nyland 1950 

Sporobolomyces Kluyver & C.B. Niel 

1924, anamorphic Sporidiobolus Nyland 

1950 (H) 

Sporidiobolales, genera incertae cedis 

Ballistosporomyces Nakase, G. Okada & 

Sugiy., in Nakase, Okada, Sugiyama, Itoh & 

Suzuki 1989 

Erythrobasidium Hamam., Sugiy. & 

Komag. 1988, anamorphic Rhodosporidium 

Banno 1967 (Y) 

Rhodotorula F.C. Harrison 1927, anamorphic 

Rhodosporidium Banno 1967 

(Y) 

Microbotryomycetes, genera incertae sedis 

Crucella Marvanová & Suberkr. 1990, 

anamorphic Camptobasidium Marvanová 

& Suberkr. 1990 (H) 

Class Pucciniomycetes R. Bauer, Begerow, 


Helicobasidiales R. Bauer, Begerow, J.P. 

Samp., M. Weiss & Oberw. 

Helicobasidiaceae P.M. Kirk 

Thanatophytum Nees 1816, anamorphic 

Helicobasidium Pat. 1885 (H) 

Tuberculina Tode ex Sacc. 1880, anamorphic 

Helicobasidium Pat. 1885 (H) 

Platygloeales R.T. Moore 

Eocronartiaceae Jülich 

Glomopsis D.M. Hend. 1961, anamorphic 

Herpobasidium Lind 1908 (H) 

Platygloeaceae Racib. 1909 

Infundibura Nag Raj & W.B. Kendr. 1981, 

anamorphic Helicogloea Pat. 1892 (H) 

Leucogloea R. Kirschner 2004 anamorphic 

Helicogloea Pat. 1892 (H) 

Uncolaceae Buriticá 

Calidion Syd. & P. Syd. 1919 (R) 

Pucciniales Clem. & Shear 

Coleosporiaceae Dietel 

?Chrysomyxa Unger 1840, anamorphic 

Peridermium (Link) J.C. Schmidt & 

Kunze 1817 (R)

Cronartiaceae Dietel 

?Cronartium Fr. 1815, anamorphic Peridermium 

(Link) J.C. Schmidt & Kunze 

1817 (R) 

Phakopsoraceae Cummins & Hirats. f. 

Aeciure Buriticá & J.F. Hennen 1994, 

anamorphic Arthuria H.S. Jacks (R) 

Macabuna Buriticá & J.F. Hennen 1994 

Malupa Y. Ono, Buriticá & J.F. Hennen 

1992 (R) 

Milesia F.B. White 1878, anamorphic 

Phakopsora Dietel 1895 but see 

Milesina Magnus 1909 (R) 

Physopella Arthur 1906, anamorphic Cerotelium 

Arthur 1906 and Phakopsora 

Dietel 1895 (R) 

Uredendo Buriticá & J.F. Hennen 1994, 

anamorphic Phakopsora Dietel 1895 

(?R) 

Uredostilbe Buriticá & J.F. Hennen 1994 

(?R) 

Phragmidiaceae Corda 

Gerwasia Racib. 1909, anamorphic Campanulospora 

Salazar-Yepes, Pardo- 

Card. & Buriticá 2007, Morispora 

Salazar-Yepes, Pardo-Card. & Buriticá 

2007 and Scutelliformis Salazar-Yepes, 

Pardo-Card. & Buriticá 2007 (R) 

Physonema Lév. 1847, anamorphic Phragmidium 

Link 1815 (?R) 

Puccineaceae Chevall. 

Caeoma Link 1809 (R) 

Roestelia Rebent. 1804, anamorphic Gymnosporangium 

R. Hedw. ex DC. 1805 

(R) 

Pucciniastraceae Gäum. ex Leppik 

Milesia F.B. White 1878, anamorphic 

Milesina Magnus 1909, but see Phakopsora 

Dietel 1895 (R) 

Peridiopsora Kamat & Sathe 1969, 

anamorphic Milesina Magnus 1909 (R) 

Pomatomyces Oerst. 1864, anamorphic 

Thekopsora Magnus 1875 (?R) 

Pucciniastrum G.H. Otth 1861, ?anamorphic 

Peridermium (Link) J.C. 

Schmidt & Kunze 1817 (R) 

Uropyxidaceae Cummins & Y. Hirats. 


Canasta A.A. Carvalho & J.F. Hennen 

2010, anamorphic Prospodium Arthur 

1907 (R) 

Pucciniales, genera incertae sedis 

Elateraecium Thirum., F. Kern & B.V. 

Patil 1966, anamorphic Hiratsukamyces 

Thirum., F. Kern & B.V. Patil 1975 (R) 

Intrapes J.F. Hennen & Figueiredo 1979 

(?R) 

Uraecium Arthur 1933 (R) 

Uredo Pers. 1801 (R) 

Septobasidiales Couch ex Donk 

Septobasidiaceae Racib. 

Johncouchia S. Hughes & Cavalc. 1983, 

anamorphic Septobasidium Pat. 1892 

(H) 

Subphylum USTILAGINOMYCOTINA R. 

Bauer, Begerow, J.P. Samp., M. Weiss & 

Oberw. 

Class Exobasidiomycetes Begerow, M. Stoll, 

R. Bauer 

Doassansiales R. Bauer & Oberw. 

Doassansiaceae R.T. Moore ex P.M. Kirk, P.F. 

Cannon & J.C. David 

Savulescuella Cif. 1959, anamorphic 

Doassansia Cornu 1883 (H) 

Entylomatales R. Bauer & Oberw. 

Entylomataceae R. Bauer & Oberw. 

Entylomella Höhn. 1924, anamorphic 

Entyloma de Bary 1874 (H) 

Tilletiopsis Derx 1948, anamorphic Entyloma 

de Bary 1874 and Melanotaenium 

de Bary 1874 (H) 

Exobasidiales Henn. 

Cladostergigma Pat. 1892 (H) 

Clinoconidium Pat. 1898 (H) 

Coniodictyum Har. & Pat. 1909 (H) 

Brachybasidiaceae Gäum. 

Meira Boekhout, Scorzetti, Gerson & Sztejnb. 

2003 (H) 

Microstromatales R. Bauer & Oberw. 

Microstromataceae Jülich 

Microstroma Niessl 1861 (H) 

Sympodiomycopsis Sugiy., Tokuoka & 

219

220 

Komag. 1991 (?Y) 

Microstromatales , genera incertae sedis 

Quambalaria J.A. Simpson 2000 (H) 

Class Ustilaginomycetes R. Bauer, Oberw. & 

Vánky 

Urocystidales R. Bauer & Oberw. 

Glomosporiaceae Cif. 

Rhombiella Liro 1939, anamorphic Thecaphora 

Fingerh. 1836 and Tothiella 

Vánky 1999 (H) 

Thecaphorella H. Scholz & I. Scholz 

1988, anamorphic Thecaphora Fingerh. 

1836 (H) 

Ustilaginales G. Winter 

Anthracoideaceae Denchev 

Crotalia Liro 1938, anamorphic Anthracoidea 

Bref. 1895 (?Y) 

Cintractiellaceae Vánky 

Naiadella Marvanová & Bandoni 1987, 

anamorphic Classicula R. Bauer, 

Begerow, Oberw. & Marvanová 2003 

(H) 

Ustilaginaceae Tul. & C. Tul. 

Pseudozyma Bandoni emend. Boekhout 

1985 (H) 

Ustilaginomycetes, orders incertae sedis 

Urocystidiales R. Bauer & Oberw. 

Urocystidaceae Begerow, R. Bauer, & Oberw. 

Paepalopsis J.G. Kühn 1882, anamorphic 

Urocystis Rabenh. ex Fuckel 1870 

Paipalopsis J.G. Kühn 1882, anamorphic 

Urocystis Rabenh. ex Fuckel 1870 (H) 

USTILAGINOMYCOTINA, order incertae 

sedis 

Malasseziales R.T. Moore 

Malassezia Baill. 1889 

USTILAGINOMYCOTINA, genera incertae 

sedis 

Botryoconis Syd. & P. Syd. 1906 (H) 

Tilletiella Zambett. 1970 (H) 

BASIDIOMYCOTA, class incertae sedis 

Wallemiomycetes Zalar, de Hoog & Schroers 

Wallemiales Zalar, de Hoog & Schroers 

Wallemiaceae R.T. Moore 


Wallemia Johan-Olsen 1887 (H) 

BASIDIOMYCOTA, genera incertae sedis 

Anguillomyces Marvanová & Bärl. 2000 

(H) 

Arcispora Marvanová & Bärl. 1998 (H) 

Bartheletica G. Arnaud, in Scheuer, R. 

Bauer, M. Lutz, Stabentheiner, Mel’nik 

& Brube 2008 (H) 

Gloeosynnema Seifert & G. Okada 1988 

(H) 

Lactydina Subram. 1978(‘1977’) (H) 

Microstella K. Ando & Tubaki 1984 (H) 

Nodulospora Marvanová & Bärl. 2000 (H) 

Pseudohelicomyces Garnica & E. Valenz. 

2000 (H) 

Trichosporonoides Haskins & J.F.T. 

Spencer 1967 (H) 

Stachycoremium Seifert 1986 (H) 

Stauriella Sivichai & E.B.G. Jones 2004 

(H) 

Discussion 

The study reveals that there are approximately 

2895 anamorphic genera names of 

which 701 genera and 95 anamorph-like genera 

are linked to teleomorphic genera names, 447 

are linked to teleomorph families, orders or 

classes, while for about 1728 (55%) genera no 

teleomorph link is known. 

The data used in this compilation are 

mostly from Index Fungorum, Species Fungorum, 

Kirk et al. (2008) and Seifert et al. (2011), 

plus publications during 2011 that were available 

before the manuscript was sent to press. 

Thus it is not inclusive and it is planned to 

improve the compilation in future issues. In 

future issues we will examine the literature 

from which connections are extracted to establish 

their basis. 

Although we have used data from peer 

reviewed published data we have no way to 

confirm the validity of the species names used 

in various phylogenetic analyses. Where generic 

types were sequenced we have stated this, 

but again it is not clear whether correct names 

have been applied to these taxa or whether they 

are strains of type material (in most cases they 

are not). 

The naming of fungi has long been 

problematic because of the dual nomenclature 

system. The system was introduced because it

was previously often impossible to link a 

sexual form with the asexual form and therefore 

there was a need to be able to name both 

forms (Shenoy et al. 2007). With the advent of 

the molecular era it is often possible to link the 

morphs, and if there is no obvious teleomorph 

it is becoming increasing feasible to establish 

the relationship of the anamorphic genus 

within the teleomorph taxonomic framework 

(Wingfield et al 2011). 

The following names were deleted from 

last years outline (Hyde et al. 2011) based on 

data in Seifert et al. (2011). 

Abgliophragma R.Y. Roy & Gujarati 

Acanthoderma Syd. & P. Syd. 

Aciculariella G. Arnaud 

Acinula Fr. 

Acrocylindrium Bonord 

Acrospira Mont. 

Acrosporium Bonord. 

Acrotheciella Koord. 

Acrothecium (Corda) Preuss 

Actinochaete Ferro 

Actinodochium Syd. 

Actinonema Pers. 

Actinospora Ingold 

Agarwalia D.P. Tiwari & P.D. Agrawal 

Albophoma Tak. Kobay., Masuma, Omura 

& Kyoto Watan. 

Aleurodomyces Buchner 

Allescheriella Henn. 

Allonema Syd. 

Alysidium Kunze 

Anthina Fr. 

Arnaudina Trotter 

Articularia Höhn. 

Asterophora Ditmar 

Asterophora Ditmar 

Astrodochium Ellis & Everh. 

Attamyces Kreisel 

Basididyma Cif. 

Basramyces Abdullah, Abdulk. & Goos 

Bhargavaella Sarj. Singh & K.S. Srivast. 

Bilboque Viégas 

Bilgramia Panwar, Purohit & Chouhan 

Bizzozeriella Speg. 

Blastoconium Cif. 

Blastostroma C.Z. Wei, Y. Harada & 

Katum. 

Botryodeorsum T.P. Devi, N. Mathur, 

Chowdhry, Jasvir Singh & O. Prakash 


Botryomyces Greco 

Brachyhelicoon G. Arnaud 

Broomeola Kuntze 

Bulleribasidium J.P. Samp., M. Weiss & R. 

Bauer 

Bulleribasidium J.P. Samp., M. Weiss & R. 

Bauer 

Bulleromyces Boekhout & Á. Fonseca 

Bulleromyces Boekhout & Á. Fonseca 

Columnophora Bubák & Vleugel 

Cicadocola Brain 

Discocolla Prill. & Delacr. 

Drechsleromyces Subram. 

Dryadomyces Gebhardt 

Eladia G. Sm. 1961 

Graphilbum H.P. Upadhyay & W.B. Kendr. 

Helicoceras Linder 

Helicosporina G. Arnaud ex Rambelli 

Helminthophora Bonord. 

Hennebertia M. Morelet 

Heydenia Fresen. 

Hormiactella Sacc. 

Hormisciopsis Sumst. 

Infrafungus Cif. 

Jainesia Gonz. Frag. & Cif. 

Keratinomyces Vanbreus. 

Loboa Cif., P.C. Azevedo, Campos & 

Carneiro 1956 

Magdalaenaea G. Arnaud 

Mammariopsis L.J. Hutchison & J. Reid 

Manoharachariomyces N.K. Rao, D.K. 

Agarwal & Kunwar 

Mastigomyces Imshen. & Kriss 

Microides H.Y.Yip & A.C. Rath 

Mucobasispora Mustafa & Abdul-Wahid 

Multicladium K.B. Deshp. & K.S. Deshp. 

Multipatina Sawada 

Myceloderma Ducomet 

Mycobacillaria Naumov 

Myropyxis Ces. ex Rabenh. 

Naranus Ts. Watan. 

Negeriella Henn. 

Obstipispora R.C. Sinclair & Morgan-Jones 

Ochroconis de Hoog & Arx 

Ommatosporella Bat., J.L. Bezerra & Poroca 

Ordus K. Ando & Tubaki 

Panchanania Subram. & N.G. Nair 

Paradactylaria Subram. & Sudha 

Paradactylella Matsush. 

Paraphialocephala Budathoki 

Periola Fr. 

Phragmodochium Höhn. 

221

222 

Plectrothrix Shear 

Pollaccia E.Bald. & Cif. 

Polyscytalina G. Arnaud 

Polysynnema Constant. & Seifert 

Polythrinciopsis J. Walker 

Poroisariopsis M. Morelet 

Pseudocladosporium Braun. 1998 

Pseudohansfordia G.R.W. Arnold 

Pseudostilbella Munt.-Cvetk. & Gómez- 

Bolea 

Pseudostracoderma A.E. Martinez & 

Godeas 

Pulvinotrichum Gamundí, Aramb. & Giaiotti 

Racodium Pers. 

Racodium Pers. 

Ramalia Bat. 1957 

Ramaraomyces N.K. Rao, Manohar. & Goos 

Ramoacrodictys G.Z. Zhao 

Ramulaspera Lindr. 

Rhabdomyces Balbiani 

Roigiella R.F. Castañeda 

Sagrahamala Subram. 

Saliastrum myrtilli (Allesch.) Kujala 

Septodochium Matsush. 

Solheimia E.F. Morris 1967 

Sphaerocolla P. Karst. 

Sphaerocybe Magrou & Marneffe 

Stenocephalum Chamuris & C.J.K. Wang 

Stephanosporium Dal Vesco 

Stevensomyces E.F. Morris & Finley 

Synnmukerjiomyces Aneja & R. Kumar 

Tretophragmia Subram. & Natarajan 

Tylomyces Cortini 

Vamsapriya Gawas & Bhat 

Vasculomyces S.F. Ashby 

Veronaella Subram. & K.R.C. Reddy 

Volutellis Clem. & Shear 

Volutellopsis Speg. 

Yunnania H.Z. Kong 

Notes 

Notes are provided on recent published 

data on anamorphic taxa. Notes are provided if 

new data is published in relation to the phylogeny, 

if changes in anamorph or teleomorph 

linkages have been revealed and if new genera 

are introduced. If well known connections are 

confirmed we may provide data from one or 

more publications, however, in many cases we 

did not cite these data as the known connections 

are unchanged. 


Acremonium Link 

Summerbell et al. (2011) generated 

sequences for members of Acremonium and 

showed that within Hypocreales, there are two 

major clusters containing multiple Acremonium 

species. One clade contains A. sclerotigenum, 

the genus Emericellopsis, and the genus 

Geosmithia as prominent elements. The second 

clade contains the genera Gliomastix sensu 

stricto and Bionectria. In addition, there were 

numerous smaller clades plus two multi-species 

clades, one containing A. strictum and related 

species plus A. curvulum and related species. 

An account of Acremonium is provided in 

Seifert et al. (2011). 

Acroconidiellina (Berk. & Broome) M.B. Ellis 

Zhang et al. (2011a) stated that Acroconidiellina 

arecae is anamorphic Acantharia 

arecae. 

Acrostalagmus Corda 

Stilbella annulata was newly combined 

in Acrostalagmus annulatus (family Plectosphaerellaceae) 

by Reblova et al. (2011). 

Alternaria Nees 

Toth et al. (2011) described a new Alternaria 

species, as A. hungarica using macro 

and micro morphological examinations and 

ITS sequence analyses. Further, it was placed 

in Pleosporaceae. An account of Alternaria is 

provided in Seifert et al. (2011). 

Antennariella Bat. & Cif. 

Chomnunti et al. (2011) suggested that 

the taxonomic placement is uncertain. 

Aquaphila Goh, K.D. Hyde & W.H. Ho 

Boonmee et al. (2011) confirmed this 

genus as anamorphic Tubeufia. 

Aspergillus P. Micheli ex Link 

Sexual reproduction was examined in 

Aspergillus nomius by Horn et al. (2011). 

Crosses between sexually compatible strains 

resulted in the formation of multiple 

nonostiolate ascocarps within stromata, which 

places the teleomorph in genus Petromyces in 

Trichocomaceae. Ascocarp and ascospore morphology 

in P. nomius were similar to that in P. 

flavus and P. parasiticus, and difference

etween telemorphs were insufficient for 

species separation. An account of Aspergillus 

is provided in Seifert et al. (2011). 

Beauveria Vuill. 

Rehner et al. (2011) carried out 

multilocus phylogeny of Beauveria based on 

partial sequences of RPB1, RPB2, TEF and the 

nuclear intergenic region, Bloc. They assessed 

diversity within the genus and evaluated 

species concepts and their taxonomic status. 

Further, they proposed types for B. bassiana 

and B. brongniartii while describing six new 

species. An account of Beauvaria is provided 

in Seifert et al. (2011). 

Chalara (Corda) Rabenh. 

Seifert et al. (2011) reported four teleomorph 

species and tentatively another twelve 

species that can be teleomorphs. 

Ciferrioxyphium Bat. & H. Maia 

The teleomorph, Aithaloderma was 

transferred to Chaetothyriaceae by Chomnunti 

et al. (2011) 

Cladobotryum Nees 

Poldmaa (2011) reported twelve new 

species of Cladobotryum/Hypomyces in 

Hypocreaceae. Ten of these were described as 

new species, including teleomorphs for two 

previously known anamorphic species; 

Cladobotryum coriolopsicola anamorphic Hypomyces 

samuelsii and Cladobotryum virescens 

anamorphic Hypomyces virescens. Also Cladobotryum 

heterosporum, C. indoafrum, and C. 

tchimbelense were described as new anamorphic 

species. 

Conidiocarpus Woron. 

Chomnunti et al. (2011) confirmed this 

genus to be anamorphic Phragmocapnias. 

Custingophora Stolk, Hennebert & Klopotek 

Species of Gondwanamyces and their 

Custingophora anamorphs were introduced 

into a new family Gondwanamycetaceae, by 

Reblova et al. (2011a). 

Cylindrocarpon Wollenw. 

Chaverri et al. (2011) showed five 

distinct highly supported clades that correspond 


to some extent with the informal Cylindrocarpon 

(anamorph) and Neonectria (telomorph) 

groups and three of which are newly 

described. The following genera were recognised: 

(1) Neonectria / Cylindrocarpon sensu 

stricto (N. coccinea-group); (2) N. rugulosagroup, 

hereafter Rugonectria gen. nov.; (3) N. 

mammoidea / veuillotiana-group, hereafter 

Thelonectria gen. nov.; (4) N. radicicola group, 

hereafter Ilyonectria gen. nov.; and (5) 

Campylocarpon. and also, results showed that 

Neonectria/Cylindrocarpon sensu stricto 

clusters with Neonectria fuckeliana, and T. 

jungneri with Thelonectria. 

Cylindrotrichum Bonord. 

Reblova et al. (2011b) introduced a new 

genus Reticulascus for Chaetosphaeria 

tulasneorum with associated Cylindrotrichum 

anamorph and placed in new family 

Reticulascaceae. 

Discosia Lib. 

Tanka et al. (2011) showed Discosia 

was not monophyletic and was separated into 

two distinct lineages. The new genus Immersidiscosia 

(family Amphisphaeriaceae) was 

erected on the basis of SSU, LSU and ITS 

nrDNA and β-tubulin gene sequences. Further, 

they stated this genus anamorphic Adisciso. 

Eriothyrium Speg. 

Wu et al. (2011) transferred the teleomorp 

to Asterinaceae and confirmed to be 

anamorphic Asterinema 

Fumiglobus D.R. Reynolds & G.S. Gilbert 

Chomnunti et al. (2011) listed the placement 

of this genus in Capnodiaceae as 

doubtful. 

Fusarium Link 

Schroers et al. (2011) reinstated the 

connection of of Fusarium buxicola with the 

bluish black and KOH+ perithecial species 

Cyanonectria buxi. The connection of F. 

buxicola is often erroneously reported as the 

anamorph of Geejayessia desmazier. 

A new species, Fusarium musae was 

described together with its teleomorph 

Gibberella musae by Hove et al. (2011). 

Scauflaire et al (2011) used a robust 

223

polyphasic approach to describe a new 

biological species, Fusarium temperatum, 

within the Gibberella fujikuroi species 

complex. 

Geosmithia Pitt 

Geosmithia morbida is a new species 

described by the Kolarik et al. (2011) and this 

is the first report of a phytopathogenic species 

in this genus and in the Bionectriaceae. G. 

morbida has been isolated only from Pityophthorus 

juglandis or from necrotic phloem 

associated with P. juglandis feeding on Juglans 

species in western USA. The telemorph is 

unknown. 

Gliomastix Guég. 

Gliomastix (Hypocreales) was revived for 

five species, G. murorum, G. polychroma, G. 

tumulicola, G. roseogrisea, and G. masseei by 

Summerbell et al. (2011) 

Heliocephala V.G. Rao, K.A. Reddy & de 

Hoog 

Abarca et al. (2011) recognized 

Heliocephala and Holubovaniella as synonymous 

and described a new species Heliocephala 

triseptata. All the species were 

positioned in Dothideomycetes and identified 

Stomiopeltis betulae (Micropeltidaceae) its 

closest relative. 

Helicoma Corda 


genus to be anamorphic Tubeufia and 

Thaxteriella Further they introduced a new 

genus Chlamydotubeufia and H. Chlamydosporum 

and H. depressispora were transered 

to a new genus. 

Helicoön Morgan 


genus to be anamorphic Tubeufiaceae. 

Helicomyces Link 

Boonmee et al. (2011) ensured that 

Helicomyces to be anamorphic Acanthostigma. 

Helicosporium Nees 

Boonmee et al. (2011) confirmed that 

Helicosporium to be anamorphic Acanthostigma 

and Tubeufia. 

224 


Holubovaniella R.F. Castañeda 

Abarca et al. (2011) placed all the 

species in Dothideomycetes and identified 

Stomiopeltis betulae (Micropeltidaceae) as 

their closest relative by analysing large subunit 

rDNA. 

Hyphopodia-like 

Suetrong et al. (2011) introduced a new 

family, Trematosphaeriaceae. Trematosphaeria 

is the type genus and Hyphopodia-like 

anamorphs were recorded. 

Immersidiscosia Kaz. Tanaka, Okane & 

Hosoya 

Discosia eucalypti deviated from 

Discosia clade and was transferred to a new 

genus, Immersidiscosia (family Amphisphaeriaceae) 

by Tanaka et al. (2011) on the basis 

of SSU, LSU and ITS nrDNA and β-tubulin 

gene sequencing. 

Leptographium Lagerb. & Melin 

Kim et al. (2011) recorded a new Leptographium 

species, L. tereforme (Ophiostomataceae) 

isolated from the bark beetle 

Hulurgus ligniperda. 

Leptoxyphium Speg. 

The teleomorph, Aithaloderma was 

transferred to Chaetothyriaceae by Chomnunti 

et al. (2011). 

Lomaantha Subram. 

Ma et al. (2011) described a new 

species Lomaantha phragmitis (anamorphic 

Pezizomycotina) and provided a key and 

synoptic table of morphological characters of 

all three Lomaantha species. 

Lolia Abdel- Aziz & Abdel- Wahab 

Phylogenetic analyses of the 28S 

ribosomal large subunit (LSU) rDNA 

sequence placed the new fungus in the family 

Lindgomycetaceae, Pleosporales. 

Mariannaea G. Arnaud ex Samson 

A new species of Mariannaea (family 

Nectriaceae) was described as M. samuelsii by 

Grafenhan et al. (2011).

Microcera Desm. 

Grafenhan et al. (2011) described a new 

species as Microcera rubra. 

Mirandina G. Arnaud ex Matsush. 

Boonmee et al. (2011) removed the 

teleomorph Taphrophila from Tubeufiaceae. 

Monilochaetes Halst. ex Harter 

Reblova et al. (2011) confirmed the 

synonymy of Monilochaetes with Dischloridium. 


Thüm. 

Chomnunti et al. (2011) stated that this 

is polyphyletic and included Coccodiniaceae. 

Phaeoxyphiella Bat. & Cif. 1963 

Chomnunti et al. (2011) confirmed 

Phaeoxyphiella to be anamorphic Capnodium. 

Polychaeton (Pers.) Lev. 


genus to be anamorphic Capnodiaceae. 

Polychaetella Speg. 

Chomnunti et al. (2011) stated that the 

status of this genus in Capnodiaceae is 

uncertain. 

Pseudotripoconidium Z.F. Yu, Ying Zhang & 

K.Q. Zhang 

A new anamorph genus Pseudotripoconidium 

was proposed by the Yu et al. 

(2011) using morphological and molecular 

analyses. It was described based on four 

isolates from ascospores of Orbilia aff. 

luteorubella. Also, this fungus differs from 

previously known Orbilia anamorphs in 

producing inversely pyramidal, unicellular 

conidia with several protuberances at their 

distal end. 

Repetophragma Subram. 

Ruiz et al. (2011) described a new 

species R. paracambrense from Mexico by the 

morphology of its conidia. 

Rhynchosporium Heinsen ex A.B. Frank 

Two new species of Rhynchosporium 

anamorphic Dermateaceae (R. commune and 


R. agropyri) were described by the Zaffarano et 

al. (2011). 

Sarcinella Sacc. 

A new species of Sarcinella (S. 

tamarindi) and its Questieriella synanamorph 

(family Englerulaceae) were reported by Hosagoudar 

& Riju (2011). 

Scolecoxyphium Cif. & Bat. 


genus to be anamorphic Capnodium and 

Scorias. 

Stachylidium Link 

Stachylidium was added in to the family 

Plectosphaerellaceae by Reblova et al. (2011) 

Sterigmatobotrys Oudem. 

Reblova & Seifert (2011) reported the 

teleomorph of Sterigmatobotrys macrocarpa 

(anamorphic Pezizomycotina) to be morphologically 

similar to the species of genera 

Carpoligna and Chaetosphaeria; but ncLSU 

phylogeny confirmed that Sterigmatobotrys is 

closely related to Carpoligna and its 

Pleurothecium anamorph and to the anamorphic 

species Pleurothecium obovoideum. 

Stilbella Lindau 

Jaklitsch & Voglmayr (2011) described 

a new telemorph Nectria eustromatica which 

has an anamorph, shares morphological traits 

with the genera Stilbella but produces nonphialidic 

macroconidia in addition to 

phialoconidia. 

Sympodiella-like 

Zhang et al. (2011a) reported Symodie- 

lla-like anamorphs from Sympoventuria. 

Trichoderma Pers. 

A new species of Trichoderma 

(anamorphic Hypocrea) (family Hypocreaceae), 

T. amazonicum, endophytic on the 

living sapwood and leaves of Hevea spp. trees 

was described by Chaverri et al. (2011). 

Trichothecium Link 

The genus Trichothecium was revised 

following the principles of unitary nomenclature 

based on the oldest valid anamorph 

225

or teleomorph name, and new combinations are 

made in Trichothecium for the tightly 

interrelated Acremonium crotocinigenum, Spicellum 

roseum, and teleomorph Leucosphaerina 

indica by Summerbell et al. (2011). 

Tripospermum Speg. 

Seifert et al. (2011) stated that 

Tripospermum to be anamorphic Triposporiopsis. 

However Chomnunti et al. (2011) 

suggested its status is uncetian. 

Tubercularia Weber ex F.H. Wigg 

Jaklitsch & Voglmayr (2011) described 

a new telemorph Nectria eustromatica with an 

anamorph that shares morphological traits with 

Tubercularia but produces non-phialidic 

macroconidia in addition to phialoconidia. 

Veronaeopsis Arzanlou & Crous 

Zhang et al. (2011a) transferred this 

genus to a new family Sympoventuriaceae. 

Zymoseptoria Quaedvlieg & Crous 

Quaedvlieg et al. (2011) described the 

new genus Zymoseptoria (anamorphic 

Mycosphaerella). 

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